Okenia gets blown apart — Bermudella, Cargoa, Ceratodoris resurrected, Naisdoris erected

A paper reclassifying Goniodorididae using DNA: Bermudella, Cargoa, and Ceratodoris are resurrected, and a new genus, Naisdoris, is described.

https://www.sciencedirect.com/science/article/pii/S1055790323002907

1. Subfamily Anculinae (hagoromo-umiushi subfamily)

Ancula (hagoromo-umiushi genus)

No major changes here. Kariya-umiushi is clearly included. (hagoromo-umiushi itself will presumably be addressed in a separate paper.)

Trapania (tsugaru-umiushi genus)

Also no major changes — but one notable sentence:

Sanger-based results of each molecular marker separately and for the concatenated alignment revealed that T. darvelli and T. reticulata are the same species

So on DNA, darvelli-umiushi and Trapania reticulata are conspecific.

2. Subfamily Goniodoridinae (nekojita-umiushi subfamily)

Goniodoridella (kotohime-umiushi genus)

Building on the kotohime piece, Okenia picoensis and Okenia felis also fall in here.

Cargoa

Contains Okenia impexa and Okenia problematica. Neither is likely to occur in Japanese waters.

Murphydoris

The genus I wrote about previously. Several undescribed species are mentioned.