Sea Slugs in Okinawa Island (East coast) 406

Hypselodoris infucata

(Rüppell & Leuckart, 1830)

Goniobranchus sinensis

(Rudman, 1985)

Chromodoris lineolata

(van Hasselt, 1824)

Hypselodoris tryoni

(Garrett, 1873)

Goniobranchus leopardus

(Rudman, 1987)

Mexichromis multituberculata

(Baba, 1953)

Phestilla melanobrachia

Bergh, 1874

Sagaminopteron multimaculatum

Ong & Gosliner, 2017

Coryphellina pseudolotos

Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022

Thecacera picta

Baba, 1972

Goniobranchus reticulatus

(Quoy & Gaimard, 1832)

Caloria indica

(Bergh, 1896)

Elysia marginata

(Pease, 1871)

Trapania naeva

Gosliner & Fahey, 2008

Goniobranchus fidelis

(Kelaart, 1858)

Goniobranchus geometricus

(Risbec, 1928)

Elysia sp. 8

Janolus flavoanulatus

Pola & Gosliner, 2019

Pteraeolidia semperi

(Bergh, 1870)

Oxynoe kylei

Krug, Berriman & Á. Valdés, 2018

Elysia pusilla

(Bergh, 1872)

Thuridilla splendens

(Baba, 1949)

Gymnodoris suggens

(Risbec, 1928)

Thuridilla gracilis

(Risbec, 1928)

Rostanga sp. 6

Tambja sp. 5

Elysia flavipunctata

M. Ichikawa, 1993

Plocamopherus pecoso

Valles & Gosliner, 2006

Hypselodoris decorata

(Risbec, 1928)

Goniobranchus fabulus

Soong & Gosliner, 2022

Unidentia aliciae

Korshunova, Mehrotra, Sp. Arnold, Lundin, Picton & Martynov, 2019

Niparaya regiscorona

(Bertsch, 1972)

Elysia asbecki

Wägele, Stemmer, Burghardt & Händeler, 2010

Doris immonda

(Risbec, 1928)

Ceratosoma gracillimum

C. Semper, 1876

Glossodoris cf. cincta

(Bergh, 1888)

Phyllodesmium poindimiei

(Risbec, 1928)

Trapania kahel

Smirnoff, Donohoo & Gosliner, 2022

Goniobranchus cf. variatus

(Risbec, 1928)

Goniobranchus coi

(Risbec, 1956)

Cyerce kikutarobabai

Hamatani, 1976

Polycera sp. 2

Hallaxa iju

Gosliner & S. Johnson, 1994

Goniobranchus hintuanensis

(Gosliner & Behrens, 1998)

Elysia spp.

Elysia sp.21

Chelidonura amoena

Bergh, 1905

Elysia tomentosa

K. R. Jensen, 1997

Elysia yaeyamana

Baba, 1936

Trapania armilla

Gosliner & Fahey, 2008

Trapania scurra

Gosliner & Fahey, 2008

Aldisa fragaria

Tibiriça, Pola & Cervera, 2017

Cadlinella ornatissima

(Risbec, 1928)

Ceratosoma trilobatum

(J. E. Gray, 1850)

Verconia simplex

(Pease, 1871)

Aeolidiopsis ransoni

Pruvot-Fol, 1956

Gymnodoris spp.

Kabeiro phasmida

Shipman & Gosliner, 2015

Hallaxa sp. 2

Haminoeidae sp. 2

Spinoaglaja orientalis

(Baba, 1949)

Thuridilla kathae

Gosliner, 1995

Sacoproteus smaragdinus

(Baba, 1949)

Trapania gibbera

Gosliner & Fahey, 2008

Trapania miltabrancha

Gosliner & Fahey, 2008

Gymnodoris impudica

(Rüppell & Leuckart, 1830)

Mexichromis aurora

(R. F. Johnson & Gosliner, 1998)

Tenellia sibogae

(Bergh, 1905)

Favorinus sp. 1

Elysia sp.14

Trapania tamaraw

Smirnoff, Donohoo & Gosliner, 2022

Odontoglaja guamensis

Rudman, 1978

Philinopsis speciosa

Pease, 1860

Elysia trisinuata

Baba, 1949

Caliphylla sp. 1

Tyrannodoris sp. 1

Okenia rhinorma

Rudman, 2007

Trapania euryeia

Gosliner & Fahey, 2008

Godiva sp. 1

Lomanotus sp. 1

Doriprismatica sp. 1

Pelagella sp. 4

Aeolidina sp. 2

Diversidoris sp. 1

Niparaya sp. 3

Kabeiro christianae

Shipman & Gosliner, 2015

Gastropteron sp. 7

Jorunna sp. 1

Ardeadoris cruenta

(Rudman, 1986)

Atys semistriatus

Pease, 1860

Sagaminopteron ornatum

Tokioka & Baba, 1964

Siphopteron tigrinum

Gosliner, 1989

Oxynoe viridis

(Pease, 1861)

Elysia sp. 5

Thuridilla livida

(Baba, 1955)

Stiliger sp. 1

Trapania aurata

Rudman, 1987

Naisdoris sp. 1

Diversidoris aurantionodulosa

Rudman, 1987

Phyllidiella nigra

(van Hasselt, 1824)

Dermatobranchus primus

Baba, 1976

Samla bicolor

(Kelaart, 1858)

Coryphellina exoptata

(Gosliner & Willan, 1991)

Favorinus tsuruganus

Baba & Abe, 1964

Placida sp. 1

Verconia sp. 1

Trapania sp. 7

Thordisa sp. 5

Eubranchus sp.10

Gastropteridae sp. 5

Haminoeidae sp. 6

Jorunna daoulasi

Innabi, Stout & Á. Valdés, 2023

Phyllidiella pustulosa

(Cuvier, 1804)

Jorunna rubescens

(Bergh, 1876)

Tubulophilinopsis pilsbryi

(Eliot, 1900)

Chromodoris colemani

Rudman, 1982

Aplysia nigrocincta

E. von Martens, 1880

Lobiger viridis

Pease, 1863

Pleurobranchus peronii

Cuvier, 1804

Hexabranchus lacer

(Cuvier, 1804)

Roboastra gracilis

(Bergh, 1877)

Halgerda willeyi

Eliot, 1904

Hallaxa indecora

(Bergh, 1905)

Goniobranchus rufomaculatus

(Pease, 1871)

Goniobranchus setoensis

(Baba, 1938)

Hypselodoris maritima

(Baba, 1949)

Verconia varians

(Pease, 1871)

Facelina bourailli

(Risbec, 1928)

Baeolidia japonica

Baba, 1933

Pleurolidia juliae

Burn, 1966

Polycera sp. 3

Trapania tigger

Smirnoff, Donohoo & Gosliner, 2022

Murphydoris sp. 4

Jorunna parva

(Baba, 1938)

Vellicolla muscaria

(A. A. Gould, 1859)

Unidentia sp. 5

Syphonota geographica

(A. Adams & Reeve, 1850)

Thuridilla vataae

(Risbec, 1928)

Colpodaspis thompsoni

G. H. Brown, 1979

Mariaglaja inornata

(Baba, 1949)

Chelidonura pallida

Risbec, 1951

Noalda sp. 2

Gastropteron sp. 3

Siphopteron ladrones

(Carlson & Hoff, 1974)

Metaruncina setoensis

(Baba, 1954)

Dolabella auricularia

([Lightfoot], 1786)

Notarchus indicus

Schweigger, 1820

Elysia sp. 1

Elysia thompsoni

K. R. Jensen, 1993

Cyerce liliuokalaniae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Costasiella sp. 1

Plocamopherus tilesii

Bergh, 1877

Tambja morosa

(Bergh, 1877)

Tayuva lilacina

(A. A. Gould, 1852)

Jorunna sp. 3

Sclerodoris rubicunda

(Baba, 1949)

Actinocyclus verrucosus

Ehrenberg, 1831

Hallaxa paulinae

Gosliner & S. Johnson, 1994

Chromodoris magnifica

(Quoy & Gaimard, 1832)

Doriprismatica atromarginata

(Cuvier, 1804)

Hypselodoris bollandi

Gosliner & R. F. Johnson, 1999

Diversidoris crocea

(Rudman, 1986)

Dendrodoris coronata

Kay & D. K. Young, 1969

Dendrodoris krusensternii

(J. E. Gray, 1850)

Dendrodoris fumata

(Rüppell & Leuckart, 1830)

Melibe megaceras

Gosliner, 1987

Myja hyotan

Martynov, Mehrotra, Chavanich, R. Nakano, Kashio, Lundin, Picton & Korshunova, 2019

Aeolidiopsis harrietae

Rudman, 1982

Thorunna sp. 4

Pelagella longicornis

Paz-Sedano, Smirnoff, Gosliner & Pola, 2023

Favorinus sp. 2

Murphydoris puncticulata

Paz-Sedano, Smirnoff, Candás, Gosliner & Pola, 2022

Niparaya sp. 4

Sohgenia sp. 1

Goniodoridella sp.16

Gymnodoris nagoensis

De Souza-Canal, R. Nakano & Á. Valdés, 2025

Jorunna sp. 7

Chelidonura hirundinina

(Quoy & Gaimard, 1833)

Sagaminopteron sp. 1

Gastropteridae sp. 1

Aplysia argus

Rüppell & Leuckart, 1830

Aplysia oculifera

A. Adams & Reeve, 1850

Phyllaplysia lafonti

(P. Fischer, 1870)

Volvatella maculata

K. R. Jensen, 2015

Elysia sp.12

Plakobranchus ocellatus

van Hasselt, 1824

Thuridilla carlsoni

Gosliner, 1995

Thuridilla flavomaculata

Gosliner, 1995

Cyerce goodheartae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Polybranchia orientalis

(Kelaart, 1858)

Sohgenia palauensis

Hamatani, 1991

Costasiella paweli

M. Ichikawa, 1993

Panderevela hyllebergi

K. R. Jensen, 2021

Kaloplocamus acutus

Baba, 1949

Naisdoris sp. 5

Trapania darvelli

Rudman, 1987

Trapania palmula

Gosliner & Fahey, 2008

Trapania vitta

Gosliner & Fahey, 2008

Aegires villosus

Farran, 1905

Gymnodoris sp. 5

Aldisa zavorensis

Tibiriçá, Pola & Cervera, 2017

Doris viridis

(Pease, 1861)

Discodoris cebuensis

Bergh, 1877

Halgerda tessellata

(Bergh, 1880)

Jorunna cf. pantherina

(Angas, 1864)

Platydoris sanguinea

Bergh, 1905

Sebadoris fragilis

(Alder & Hancock, 1864)

Ceratosoma tenue

Abraham, 1876

Goniobranchus kuniei

(Pruvot-Fol, 1930)

Goniobranchus preciosus

(Kelaart, 1858)

Goniobranchus verrieri

(Crosse, 1875)

Hypselodoris bullockii

(Collingwood, 1881)

Hypselodoris whitei

(A. Adams & Reeve, 1850)

Mexichromis mariei

(Crosse, 1872)

Ardeadoris angustolutea

(Rudman, 1990)

Verconia norba

(Er. Marcus & Ev. Marcus, 1970)

Thorunna australis

(Risbec, 1928)

Thorunna daniellae

(Kay & D. K. Young, 1969)

Dendrodoris nigra

(W. Stimpson, 1855)

Phyllidia coelestis

Bergh, 1905

Phyllidia exquisita

Brunckhorst, 1993

Madrella ferruginosa

Alder & Hancock, 1864

Lomanotus vermiformis

Eliot, 1908

Pruvotfolia sp. 3

Bulbaeolidia alba

(Risbec, 1928)

Tenellia sp. 5

Phestilla lugubris

(Bergh, 1870)

Siphopteron makisig

Ong & Gosliner, 2017

Goniobranchus sp. 6

Chromodoris sp. 3

Acteocina sp.

Trapania kamagong

Smirnoff, Donohoo & Gosliner, 2022

Murphydoris maracabranchia

Paz-Sedano, Smirnoff, Candás, Gosliner & Pola, 2022

Tenellia sp.10

Facelina sp. 3

Gastropteridae sp. 3

Novaeolidiella janae

Carmona, Martín-Hervás, Pola, Gosliner & Cervera, 2026

Baeolidia sp. 1

Tenellia puti

A. Y. Kim & Gosliner, 2024

Tenellia sp.67

Cyerce katiae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Armina sp. 1

Limapontiidae spp.

Plocamopherus sp. 3

Mariaglaja tsurugensis

(Baba & Abe, 1959)

Haminoeidae sp. 7

Facelinidae sp. 6

Costasiella sp. 6

Gymnodoris plebeia

(Bergh, 1877)

Gymnodoris sp.31

Micromelo guamensis

(Quoy & Gaimard, 1825)

Trapania tatsulok

Smirnoff, Donohoo & Gosliner, 2022

Gastropteron sp. 8

Biuve sp. 1

Tylodina sp. 1

Tritonia bollandi

V. G. Smith & Gosliner, 2003

Hypselodoris purpureomaculosa

Hamatani, 1995

Bornella stellifera

(A. Adams & Reeve, 1848)

Roxaniella multistriata

(Schepman, 1913)

Lamprohaminoea ovalis

(Pease, 1868)

Chelidonura sp. 1

Noalda sp. 1

Gastropteron minutum

Ong & Gosliner, 2017

Sagaminopteron bilealbum

Carlson & Hoff, 1973

Sagaminopteron psychedelicum

Carlson & Hoff, 1974

Aplysia sagamiana

Baba, 1949

Petalifera punctulata

(Tapparone Canefri, 1874)

Stylocheilus striatus

(Quoy & Gaimard, 1832)

Julia japonica

Kuroda & Habe, 1951

Bosellia sp. 1

Elysia obtusa

Baba, 1938

Elysia mercieri

(Pruvot-Fol, 1930)

Elysia nealae

Ostergaard, 1955

Costasiella kuroshimae

M. Ichikawa, 1993

Costasiella usagi

M. Ichikawa, 1993

Placida kevinleei

McCarthy, Krug & Á. Valdés, 2017

Stiliger aureomarginatus

K. R. Jensen, 1993

Polycera sp. 7

Polycera risbeci

Odhner, 1941

Kaloplocamus dokte

Vallès & Gosliner, 2006

Nembrotha kubaryana

Bergh, 1877

Martadoris limaciformis

(Eliot, 1908)

Bermudella japonica

(Baba, 1949)

Aegires exeches

Fahey & Gosliner, 2004

Notodoris citrina

Bergh, 1875

Gymnodoris citrina

(Bergh, 1877)

Gymnodoris inornata

(Bergh, 1880)

Gymnodoris okinawae

Baba, 1936

Gymnodoris feyae

De Souza-Canal & Valdés, 2025

Gymnodoris striata

(Eliot, 1908)

Aldisa albatrossae

Elwood, Á. Valdés & Gosliner, 2000

Doriopsis pecten

(Collingwood, 1881)

Atagema intecta

(Kelaart, 1858)

Atagema spongiosa

(Kelaart, 1858)

Carminodoris bifurcata

Baba, 1993

Halgerda elegans

Bergh, 1905

Paradoris tsurugensis

Baba, 1986

Platydoris inornata

Dorgan, Á. Valdés & Gosliner, 2002

Thordisa luteola

J. M. Chan & Gosliner, 2007

Miamira miamirana

(Bergh, 1875)

Ceratosoma sp. 1

Goniobranchus albonares

(Rudman, 1990)

Goniobranchus rubrocornutus

(Rudman, 1985)

Goniobranchus sp.16

Mexichromis pusilla

(Bergh, 1874)

Ardeadoris averni

(Rudman, 1985)

Glossodoris buko

S. B. Matsuda & Gosliner, 2018

Ardeadoris symmetrica

(Rudman, 1990)

Ardeadoris tomsmithi

(Bertsch & Gosliner, 1989)

Hypselodoris iacula

Gosliner & R. F. Johnson, 1999

Hypselodoris krakatoa

Gosliner & R. F. Johnson, 1999

Hypselodoris zephyra

Gosliner & R. F. Johnson, 1999

Diversidoris flava

(Eliot, 1904)

Verconia romeri

(Risbec, 1928)

Thorunna punicea

(Rudman, 1995)

Dendrodoris arborescens

(Collingwood, 1881)

Phyllidia ocellata

Cuvier, 1804

Phyllidia picta

Pruvot-Fol, 1957

Phyllidia varicosa

Lamarck, 1801

Phyllidia larryi

(Brunckhorst, 1993)

Dermatobranchus rodmani

Gosliner & Fahey, 2011

Pinufius rebus

Er. Marcus & Ev. Marcus, 1960

Janolus tricellariodes

Pola & Gosliner, 2019

Marianina rosea

(Pruvot-Fol, 1930)

Tritonia sp. 1

Doto cervicenigra

Ortea & Bouchet, 1989

Coryphellina delicata

(Gosliner & Willan, 1991)

Coryphellina sp. 2

Cratena lineata

(Eliot, 1905)

Favorinus japonicus

Baba, 1949

Favorinus mirabilis

Baba, 1955

Godiva rachelae

Rudman, 1980

Herviella albida

Baba, 1966

Noumeaella isa

Ev. Marcus & Er. Marcus, 1970

Phyllodesmium iriomotense

Baba, 1991

Phyllodesmium longicirrum

(Bergh, 1905)

Phyllodesmium macphersonae

(Burn, 1962)

Pruvotfolia rhodopos

(Yonow, 2000)

Baeolidia moebii

Bergh, 1888

Cerberilla affinis

Bergh, 1888

Tenellia diversicolor

(Baba, 1975)

Tenellia sp. 2

Atys sp. 1

Janolus savinkini

Martynov & Korshunova, 2012

Gymnodoris sp. 9

Tergiposacca longicerata

Cella, Carmona, Ekimova, Chichvarkhin, Schepetov & Gosliner, 2016

Arminidae spp.

Tenellia sp.27

Tenellia nakapila

A. Y. Kim & Gosliner, 2024

Eubranchus sp. 3

Phyllidiidae spp.

Aeolidina spp.

Kabeiro sp. 1

Gymnodoris inariensis

Hamatani & Osumi, 2004

Atys sp. 3

Abronica sp. 1

Phyllodesmium sp. 2

Anteaeolidiella takanosimensis

(Baba, 1930)

Jorunna sp. 2

Tenellia sp. 8

Thordisa sp. 4

Tenellia sp.25

Atagema sp. 2

Trapania sp.13

Gymnodoris sp.14

Baeolidia variabilis

Carmona, Pola, Gosliner & Cervera, 2014

Chromodorididae spp.

Gymnodoris sp.15

Gymnodoris sp.16

Tenellia sp.77

Facelinidae sp. 2

Gymnodoris sp.18

Atagema sp. 4

Gymnodoris sp.20

Favorinus sp. 7

Tenellia sp.55

Phidiana sp. 2

Costasiella sp. 4

Gastropteridae sp. 6

Tenellia sp.72

Gymnodoris sp.23

Unidentia sandramillenae

Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017

Polybranchia spp.

Produnga rubropunctata

(Edmunds, 1969)

Siphopteron dumbo

Ong & Gosliner, 2017

Thecacera sp.12

Discodorididae sp. 1

Niparaya sp. 1

Facelina sp. 4

Hermaea sp. 4

Elysia minima

M. Ichikawa, 1993

Vellicolla amphorella

(A. Adams, 1862)

Gastropteridae spp.

Aplysia sp. 1

Gymnodoris bicolor

(Alder & Hancock, 1864)

Ercolania sp. 8

Doris sp. 2

Discodorididae sp.15

Mexichromis sp. 1

Hypselodoris spp.

Julia mishimaensis

Kawaguti & Yamasu, 1982

Edenttellina pseudochloris

(Kay, 1964)

Gymnodoris fleuroti

De Souza-Canal & Valdés, 2025

Elysia sp.38

Ercolania annelyleorum

Wägele, Stemmer, Burghardt & Händeler, 2010

Mourgona sp. 1