Sea Slugs in Okinawa Island (Chatan and Southern area) 425

Hypselodoris bullockii

(Collingwood, 1881)

Elysia pusilla

(Bergh, 1872)

Chromodoris sp. 3

Siphopteron ladrones

(Carlson & Hoff, 1974)

Tyrannodoris luteolineata

(Baba, 1936)

Bornella anguilla

S. Johnson, 1984

Sagaminopteron ornatum

Tokioka & Baba, 1964

Gymnodoris okinawae

Baba, 1936

Halgerda tessellata

(Bergh, 1880)

Paradoris sp. 2

Elysia asbecki

Wägele, Stemmer, Burghardt & Händeler, 2010

Odontoglaja guamensis

Rudman, 1978

Aplysia nigrocincta

E. von Martens, 1880

Tambja morosa

(Bergh, 1877)

Hypselodoris decorata

(Risbec, 1928)

Facelina bourailli

(Risbec, 1928)

Godiva sp. 1

Phyllodesmium longicirrum

(Bergh, 1905)

Gymnodoris spp.

Phyllodesmium sp. 8

Lamprohaminoea ovalis

(Pease, 1868)

Siphopteron nigromarginatum

Gosliner, 1989

Siphopteron tigrinum

Gosliner, 1989

Elysia sp. 7

Plakobranchus ocellatus

van Hasselt, 1824

Notodoris citrina

Bergh, 1875

Cadlinella ornatissima

(Risbec, 1928)

Ceratosoma gracillimum

C. Semper, 1876

Hypselodoris whitei

(A. Adams & Reeve, 1850)

Dermatobranchus fortunatus

(Bergh, 1888)

Janolus mirabilis

Baba & Abe, 1970

Janolus tricellariodes

Pola & Gosliner, 2019

Elysia nigropunctata

(Pease, 1871)

Phyllidiella pustulosa

(Cuvier, 1804)

Sagaminopteron psychedelicum

Carlson & Hoff, 1974

Elysia tomentosa

K. R. Jensen, 1997

Polycera sp. 1

Nembrotha livingstonei

J. K. Allan, 1933

Trapania naeva

Gosliner & Fahey, 2008

Goniobranchus reticulatus

(Quoy & Gaimard, 1832)

Phyllidia elegans

Bergh, 1869

Janolus toyamensis

Baba & Abe, 1970

Samla bicolor

(Kelaart, 1858)

Cratena sp.11

Cratena sp. 1

Phyllodesmium koehleri

Burghardt, Schrödl & Wägele, 2008

Bulbaeolidia alba

(Risbec, 1928)

Ercolania sp. 1

Peltodoris murrea

(Abraham, 1877)

Limapontiidae spp.

Martadoris sp. 3

Tomoberthella martensi

(Pilsbry, 1896)

Jorunna rubescens

(Bergh, 1876)

Tritoniopsis elegans

(Audouin, 1826)

Colpodaspis thompsoni

G. H. Brown, 1979

Chelidonura amoena

Bergh, 1905

Chelidonura pallida

Risbec, 1951

Odontoglaja sp. 1

Elysia flavipunctata

M. Ichikawa, 1993

Thuridilla carlsoni

Gosliner, 1995

Polybranchia orientalis

(Kelaart, 1858)

Nembrotha chamberlaini

Gosliner & Behrens, 1997

Notodoris gardineri

Eliot, 1906

Gymnodoris suggens

(Risbec, 1928)

Glossodoris cf. cincta

(Bergh, 1888)

Phyllidiidae sp. 1

Phyllidiella annulata

(J. E. Gray, 1853)

Tenellia sp.11

Phanerophthalmus lentigines

Austin, Gosliner & Malaquias, 2018

Hallaxa sp. 2

Doto sp.23

Haminoeidae sp.12

Tubulophilinopsis pilsbryi

(Eliot, 1900)

Chromodoris colemani

Rudman, 1982

Diniatys dentifer

(A. Adams, 1850)

Ilbia mariana

Hoff & Carlson, 1990

Elysia rufescens

(Pease, 1871)

Elysia thompsoni

K. R. Jensen, 1993

Thuridilla flavomaculata

Gosliner, 1995

Thuridilla kathae

Gosliner, 1995

Costasiella paweli

M. Ichikawa, 1993

Polycera japonica

Baba, 1949

Nembrotha sp. 1

Tambja sagamiana

(Baba, 1955)

Gymnodoris sp. 3

Gymnodoris sp. 5

Halgerda albocristata

Gosliner & Fahey, 1998

Jorunna funebris

(Kelaart, 1858)

Paradoris sp. 1

Actinocyclus verrucosus

Ehrenberg, 1831

Goniobranchus geometricus

(Risbec, 1928)

Phyllidia coelestis

Bergh, 1905

Phyllidia ocellata

Cuvier, 1804

Phyllidia varicosa

Lamarck, 1801

Janolus incrustans

Pola & Gosliner, 2019

Melibe engeli

Risbec, 1937

Coryphellina delicata

(Gosliner & Willan, 1991)

Launsina tanyae

Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026

Costasiella sp. 5

Thorunna sp. 4

Doriprismatica sp. 1

Bullina lineata

(J. E. Gray, 1825)

Avaldesia albomacula

(J. M. Chan & Gosliner, 2007)

Costasiella sp. 4

Hermaea wrangeliae

(M. Ichikawa, 1993)

Goniobranchus coi

(Risbec, 1956)

Tubulophilinopsis gardineri

(Eliot, 1903)

Bornella stellifera

(A. Adams & Reeve, 1848)

Smaragdinella calyculata

(Broderip & G. B. Sowerby I, 1829)

Sagaminopteron sp. 1

Siphopteron brunneomarginatum

(Carlson & Hoff, 1974)

Ascobulla japonica

(Hamatani, 1969)

Elysia obtusa

Baba, 1938

Thuridilla gracilis

(Risbec, 1928)

Thuridilla splendens

(Baba, 1949)

Cyerce basi

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Mourgona sp. 2

Sacoproteus smaragdinus

(Baba, 1949)

Aplysiopsis orientalis

(Baba, 1949)

Hexabranchus lacer

(Cuvier, 1804)

Kaloplocamus dokte

Vallès & Gosliner, 2006

Plocamopherus imperialis

Angas, 1864

Roboastra gracilis

(Bergh, 1877)

Martadoris limaciformis

(Eliot, 1908)

Pelagella joubini

(Risbec, 1928)

Aegires incusus

Fahey & Gosliner, 2004

Halgerda elegans

Bergh, 1905

Hallaxa paulinae

Gosliner & S. Johnson, 1994

Miamira sinuata

(van Hasselt, 1824)

Goniobranchus fidelis

(Kelaart, 1858)

Goniobranchus rufomaculatus

(Pease, 1871)

Verconia simplex

(Pease, 1871)

Phyllidia exquisita

Brunckhorst, 1993

Phyllidia picta

Pruvot-Fol, 1957

Phyllidiella granulata

Brunckhorst, 1993

Dermatobranchus striatus

van Hasselt, 1824

Facelina sp. 1

Phidiana salaamica

Rudman, 1980

Phyllodesmium kabiranum

Baba, 1991

Tenellia diversicolor

(Baba, 1975)

Goniobranchus sp. 6

Tergiposacca longicerata

Cella, Carmona, Ekimova, Chichvarkhin, Schepetov & Gosliner, 2016

Pelagella longicornis

Paz-Sedano, Smirnoff, Gosliner & Pola, 2023

Pelagella sp. 6

Aplustrum amplustre

(Linnaeus, 1758)

Doridoidea spp.

Coryphellina flamma

Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022

Siphopteron dumbo

Ong & Gosliner, 2017

Tritoniopsis sp. 1

Ercolania sp. 7

Ercolania sp.10

Bursatella ocelligera

(Bergh, 1902)

Mourgona sp. 4

Ercolania sp. 9

Thuridilla vataae

(Risbec, 1928)

Ardeadoris cruenta

(Rudman, 1986)

Reticulidia fungia

Brunckhorst & Gosliner, 1993

Phanerophthalmus perpallidus

Risbec, 1928

Niparaya regiscorona

(Bertsch, 1972)

Mariaglaja inornata

(Baba, 1949)

Gastropteron minutum

Ong & Gosliner, 2017

Dolabrifera dolabrifera

(Rang, 1828)

Julia zebra

Kawaguchi, 1981

Oxynoe kabirensis

Hamatani, 1980

Oxynoe kylei

Krug, Berriman & Á. Valdés, 2018

Oxynoe viridis

(Pease, 1861)

Bosellia sp. 1

Elysia marginata

(Pease, 1871)

Elysia sp. 5

Elysia sp.12

Elysia yaeyamana

Baba, 1936

Cyerce bourbonica

Yonow, 2012

Cyerce blackburnae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Cyerce liliuokalaniae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Sohgenia palauensis

Hamatani, 1991

Ercolania subviridis

(Baba, 1959)

Hermaea noto

(Baba, 1959)

Pleurobranchus forskalii

Rüppell & Leuckart, 1828

Pleurobranchus peronii

Cuvier, 1804

Plocamopherus pecoso

Valles & Gosliner, 2006

Trapania euryeia

Gosliner & Fahey, 2008

Aegires flores

Fahey & Gosliner, 2004

Aegires villosus

Farran, 1905

Gymnodoris sp. 7

Gymnodoris sp.33

Ardeadoris egretta

Rudman, 1984

Miamira miamirana

(Bergh, 1875)

Chromodoris aspersa

(A. A. Gould, 1852)

Goniobranchus setoensis

(Baba, 1938)

Mexichromis pusilla

(Bergh, 1874)

Glossodoris buko

S. B. Matsuda & Gosliner, 2018

Glossodoris rufomarginata

(Bergh, 1890)

Hypselodoris sp.18

Hypselodoris maculosa

(Pease, 1871)

Hypselodoris zephyra

Gosliner & R. F. Johnson, 1999

Ardeadoris angustolutea

(Rudman, 1990)

Verconia romeri

(Risbec, 1928)

Verconia varians

(Pease, 1871)

Mexichromis trilineata

(A. Adams & Reeve, 1850)

Thorunna halourga

R. F. Johnson & Gosliner, 2001

Phyllidiopsis xishaensis

(G.-Y. Lin, 1983)

Dermatobranchus rodmani

Gosliner & Fahey, 2011

Madrella ferruginosa

Alder & Hancock, 1864

Janolus sp. 2

Caloria indica

(Bergh, 1896)

Cratena lineata

(Eliot, 1905)

Herviella affinis

Baba, 1960

Phyllodesmium crypticum

Rudman, 1981

Phyllodesmium macphersonae

(Burn, 1962)

Aeolidiopsis ransoni

Pruvot-Fol, 1956

Baeolidia japonica

Baba, 1933

Phestilla minor

Rudman, 1981

Placida sp. 1

Pteraeolidia semperi

(Bergh, 1870)

Phyllodesmium sp. 1

Trapania sp. 7

Phyllodesmium acanthorhinum

E. Moore & Gosliner, 2014

Pruvotfolia sp. 1

Phyllidiidae spp.

Ercolania sp. 2

Tritoniopsis cf. frydis

Lamprohaminoea evelinae

Oskars & Malaquias, 2020

Elysia sp.14

Limenandra confusa

Carmona, Pola, Gosliner & Cervera, 2014

Kaloplocamus sp. 4

Gymnodoris sp.17

Gymnodoris sp.19

Phanerophthalmus albotriangulatus

Austin, Gosliner & Malaquias, 2018

Phestilla viei

Mehrotra, Caballer & Chavanich, 2020

Polybranchia spp.

Niparaya sp. 6

Janolus sp.18

Doriopsilla sp. 2

Mourgona osumi

Hamatani, 1994

Gastropteron multo

Ong & Gosliner, 2017

Edenttellina pseudochloris

(Kay, 1964)

Gymnodoris nagoensis

De Souza-Canal, R. Nakano & Á. Valdés, 2025

Unknown Species

Syphonota geographica

(A. Adams & Reeve, 1850)

Tylodina sp. 1

Jorunna sp. 1

Babakina indopacifica

Gosliner, Gonzalez-Duarte & Cervera, 2007

Atys semistriatus

Pease, 1860

Hermania orca

(Gosliner, 1988)

Chelidonura hirundinina

(Quoy & Gaimard, 1833)

Tubulophilinopsis lineolata

(H. Adams & A. Adams, 1854)

Gastropteron sp. 3

Metaruncina setoensis

(Baba, 1954)

Dolabella auricularia

([Lightfoot], 1786)

Notarchus indicus

Schweigger, 1820

Phyllaplysia lafonti

(P. Fischer, 1870)

Petalifera punctulata

(Tapparone Canefri, 1874)

Petalifera ramosa

Baba, 1959

Stylocheilus striatus

(Quoy & Gaimard, 1832)

Volvatella ayakii

Hamatani, 1972

Elysia lobata

A. A. Gould, 1852

Elysia sp. 8

Cyerce kikutarobabai

Hamatani, 1976

Cyerce goodheartae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Costasiella kuroshimae

M. Ichikawa, 1993

Costasiella sp. 1

Costasiella usagi

M. Ichikawa, 1993

Pleurehdera sp. 1

Pleurobranchus albiguttatus

(Bergh, 1905)

Pleurobranchus mamillatus

Quoy & Gaimard, 1832

Nembrotha lineolata

Bergh, 1905

Ceratodoris lambat

(Gosliner, 2004)

Trapania gibbera

Gosliner & Fahey, 2008

Trapania palmula

Gosliner & Fahey, 2008

Naisdoris sp. 1

Aegires exeches

Fahey & Gosliner, 2004

Aegires petalis

Fahey & Gosliner, 2004

Gymnodoris ceylonica

(Kelaart, 1858)

Gymnodoris citrina

(Bergh, 1877)

Gymnodoris inornata

(Bergh, 1880)

Gymnodoris feyae

De Souza-Canal & Valdés, 2025

Gymnodoris striata

(Eliot, 1908)

Gymnodoris subornata

Baba, 1960

Aldisa albatrossae

Elwood, Á. Valdés & Gosliner, 2000

Aldisa pikokai

Bertsch & S. Johnson, 1982

Aldisa zavorensis

Tibiriçá, Pola & Cervera, 2017

Doris immonda

(Risbec, 1928)

Doriopsis pecten

(Collingwood, 1881)

Atagema spongiosa

(Kelaart, 1858)

Carminodoris bifurcata

Baba, 1993

Carminodoris estrelyado

(Gosliner & Behrens, 1998)

Carminodoris grandiflora

(Pease, 1860)

Discodoris cebuensis

Bergh, 1877

Halgerda brunneomaculata

Carlson & Hoff, 1993

Halgerda diaphana

Fahey & Gosliner, 1999

Platydoris scabra

(Cuvier, 1804)

Hallaxa fuscescens

(Pease, 1871)

Hallaxa hileenae

Gosliner & S. Johnson, 1994

Hallaxa iju

Gosliner & S. Johnson, 1994

Hallaxa indecora

(Bergh, 1905)

Ceratosoma palliolatum

Rudman, 1988

Ceratosoma sp. 1

Ceratosoma tenue

Abraham, 1876

Chromodoris sp.14

Chromodoris annae

Bergh, 1877

Goniobranchus decorus

(Pease, 1860)

Goniobranchus kuniei

(Pruvot-Fol, 1930)

Goniobranchus sinensis

(Rudman, 1985)

Chromodoris strigata

Rudman, 1982

Goniobranchus verrieri

(Crosse, 1875)

Chromodoris willani

Rudman, 1982

Ardeadoris electra

(Rudman, 1990)

Glossodoris hikuerensis

(Pruvot-Fol, 1954)

Ardeadoris symmetrica

(Rudman, 1990)

Hypselodoris apolegma

(Yonow, 2001)

Mexichromis multituberculata

(Baba, 1953)

Verconia alboannulata

(Rudman, 1986)

Verconia sp. 2

Hypselodoris tryoni

(Garrett, 1873)

Thorunna daniellae

(Kay & D. K. Young, 1969)

Thorunna florens

(Baba, 1949)

Thorunna purpuropedis

Rudman & S. Johnson, 1985

Phyllidiella nigra

(van Hasselt, 1824)

Phyllidiopsis cardinalis

Bergh, 1876

Phyllidiopsis fissurata

Brunckhorst, 1993

Phyllidiopsis krempfi

Pruvot-Fol, 1957

Phyllidiopsis sphingis

Brunckhorst, 1993

Dermatobranchus caeruleomaculatus

Gosliner & Fahey, 2011

Dermatobranchus semilunus

Gosliner & Fahey, 2011

Dermatobranchus fasciatus

Gosliner & Fahey, 2011

Pinufius rebus

Er. Marcus & Ev. Marcus, 1960

Madrella gloriosa

Baba, 1949

Marioniopsis arborescens

(Bergh, 1890)

Tritonia sp. 1

Phylliroe bucephala

Lamarck, 1816

Doto rosacea

Baba, 1949

Samla riwo

(Gosliner & Willan, 1991)

Eubranchus sp. 1

Myja hyotan

Martynov, Mehrotra, Chavanich, R. Nakano, Kashio, Lundin, Picton & Korshunova, 2019

Cratena simba

Edmunds, 1970

Favorinus japonicus

Baba, 1949

Moridilla brockii

Bergh, 1888

Phyllodesmium iriomotense

Baba, 1991

Phyllodesmium lizardense

Burghardt, Schrödl & Wägele, 2008

Phyllodesmium magnum

Rudman, 1991

Phyllodesmium poindimiei

(Risbec, 1928)

Cerberilla sp. 2

Tenellia sp. 5

Phestilla lugubris

(Bergh, 1870)

Hermaea sp. 1

Cadlinella subornatissima

Baba, 1996

Nakamigawaia nakanoae

Hellem & Malaquias, 2022

Lomanotus sp. 1

Nembrotha sp. 3

Janolus savinkini

Martynov & Korshunova, 2012

Gymnodoris sp. 6

Gymnodoris mariefranceae

De Souza-Canal & Valdés, 2025

Murphydoris sp. 4

Tenellia sp.27

Tenellia sp.51

Tenellia sp. 9

Micromelo barbarae

Feliciano, Malaquias, Stout, Brenzinger, Gosliner & Á. Valdés, 2021

Eubranchus sp. 3

Hypselodoris lacuna

Gosliner & R. F. Johnson, 2018

Tenellia sp.10

Hallaxa sp. 1

Aeolidina spp.

Niparaya sp. 3

Pruvotfolia sp. 2

Facelinidae sp. 1

Bornella hermanni

Angas, 1864

Anteaeolidiella takanosimensis

(Baba, 1930)

Tenellia sp. 8

Coryphellina sp. 1

Siphopteron sp.12

Gymnodoris sp.14

Favorinus sp. 2

Baeolidia variabilis

Carmona, Pola, Gosliner & Cervera, 2014

Gymnodoris sp.16

Tenellia sp.77

Tenellia sp.30

Discodorididae sp. 5

Cyerce katiae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Favorinus sp. 8

Elysia sp.23

Phidiana sp. 2

Sagaminopteron multimaculatum

Ong & Gosliner, 2017

Elysia sp.26

Dendrodoris sp. 3

Aeolidina sp.11

Abronica sp. 5

Tenellia sp.23

Placida daguilarensis

K. Jensen, 1990

Stiliger sp.10

Noumeaella sp. 4

Vellicolla muscaria

(A. A. Gould, 1859)

Samla sp. 2

Facelina sp. 4

Hermaea sp. 4

Gymnodoris sp.27

Tenellia sp.15

Costasiella sp. 6

Eubranchus sp.28

Gymnodoris sp.28

Vellicolla amphorella

(A. Adams, 1862)

Placida barackobamai

McCarthy, Krug & Á. Valdés, 2017

Dermatobranchus sp. 5

Aplysia sp. 1

Costasiella sp. 7

Cratena sp.10

Runcinida sp. 3

Gymnodoris sp.31

Gymnodoris bicolor

(Alder & Hancock, 1864)

Ercolania sp. 8

Atys naucum

(Linnaeus, 1758)

Aplysiidae spp.

Tenellia sp.95

Thordisa sp.13

Avaldesia spp.

Nembrothinae spp.

Hypselodoris spp.

Costasiella rubrolineata

M. Ichikawa, 1993

Gymnodoris fassioae

De Souza-Canal & Valdés, 2025

Doridoidea sp.15

Coryphellina pseudolotos

Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022

Gymnodoris sp.43

Elysia sp.40

Facelina sp. 8

Ercolania annelyleorum

Wägele, Stemmer, Burghardt & Händeler, 2010

Volvatella viridis

Hamatani, 1976