Sea Slugs in Ishigaki and Yaeyama 514

Thecacera pacifica

(Bergh, 1884)

Chromodoris willani

Rudman, 1982

Mariaglaja mandroroa

(Gosliner, 2011)

Cratena sp. 1

Pruvotfolia sp. 1

Chromodoris colemani

Rudman, 1982

Elysia asbecki

Wägele, Stemmer, Burghardt & Händeler, 2010

Ceratodoris kondoi

(Hamatani, 2001)

Halgerda sp. 2

Goniobranchus fidelis

(Kelaart, 1858)

Chromodoris sp. 3

Gymnodoris spp.

Phyllidiella pustulosa

(Cuvier, 1804)

Costasiella paweli

M. Ichikawa, 1993

Nembrotha kubaryana

Bergh, 1877

Hypselodoris decorata

(Risbec, 1928)

Cratena sp.11

Chelidonura amoena

Bergh, 1905

Sagaminopteron psychedelicum

Carlson & Hoff, 1974

Aplysia nigrocincta

E. von Martens, 1880

Costasiella sp. 1

Costasiella usagi

M. Ichikawa, 1993

Roboastra gracilis

(Bergh, 1877)

Jorunna funebris

(Kelaart, 1858)

Cadlinella ornatissima

(Risbec, 1928)

Goniobranchus geometricus

(Risbec, 1928)

Phyllidia ocellata

Cuvier, 1804

Facelina bourailli

(Risbec, 1928)

Siphopteron dumbo

Ong & Gosliner, 2017

Bornella anguilla

S. Johnson, 1984

Thuridilla splendens

(Baba, 1949)

Cyerce bourbonica

Yonow, 2012

Polycera japonica

Baba, 1949

Nembrotha cristata

Bergh, 1877

Halgerda brunneomaculata

Carlson & Hoff, 1993

Goniobranchus kuniei

(Pruvot-Fol, 1930)

Verconia romeri

(Risbec, 1928)

Tenellia sp.11

Limenandra confusa

Carmona, Pola, Gosliner & Cervera, 2014

Elysia aowthai

Mehrotra, Caballer, C. M. Scott, Sp. Arnold, Monchanin & Chavanich, 2020

Siphopteron ladrones

(Carlson & Hoff, 1974)

Elysia mercieri

(Pruvot-Fol, 1930)

Costasiella fridae

Fernández-Simón & Moles, 2023

Halgerda tessellata

(Bergh, 1880)

Goniobranchus setoensis

(Baba, 1938)

Hypselodoris tryoni

(Garrett, 1873)

Phyllidia coelestis

Bergh, 1905

Samla riwo

(Gosliner & Willan, 1991)

Launsina tanyae

Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026

Godiva sp. 1

Elysia spp.

Pteraeolidia semperi

(Bergh, 1870)

Coryphellina flamma

Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022

Gastropteron sp. 3

Sagaminopteron ornatum

Tokioka & Baba, 1964

Siphopteron tigrinum

Gosliner, 1989

Tyrannodoris luteolineata

(Baba, 1936)

Goniobranchus albonares

(Rudman, 1990)

Thorunna daniellae

(Kay & D. K. Young, 1969)

Dendrodoris nigra

(W. Stimpson, 1855)

Janolus incrustans

Pola & Gosliner, 2019

Samla bicolor

(Kelaart, 1858)

Tergiposacca longicerata

Cella, Carmona, Ekimova, Chichvarkhin, Schepetov & Gosliner, 2016

Hallaxa sp. 1

Dermatobranchus diagonalis

Gosliner & Fahey, 2011

Janolus sp. 7

Hypselodoris skyleri

Gosliner & R. F. Johnson, 2018

Gymnodoris sp.21

Thuridilla vataae

(Risbec, 1928)

Ardeadoris cruenta

(Rudman, 1986)

Lamprohaminoea cymbalum

(Quoy & Gaimard, 1833)

Lamprohaminoea ovalis

(Pease, 1868)

Siphopteron brunneomarginatum

(Carlson & Hoff, 1974)

Oxynoe viridis

(Pease, 1861)

Elysia obtusa

Baba, 1938

Thuridilla flavomaculata

Gosliner, 1995

Pleurobranchus peronii

Cuvier, 1804

Martadoris limaciformis

(Eliot, 1908)

Tambja morosa

(Bergh, 1877)

Aegires flores

Fahey & Gosliner, 2004

Notodoris serenae

Gosliner & Behrens, 1997

Halgerda diaphana

Fahey & Gosliner, 1999

Goniobranchus rufomaculatus

(Pease, 1871)

Glossodoris cf. cincta

(Bergh, 1888)

Hypselodoris sp.18

Verconia simplex

(Pease, 1871)

Dermatobranchus fortunatus

(Bergh, 1888)

Dermatobranchus striatus

van Hasselt, 1824

Caloria indica

(Bergh, 1896)

Noumeaella rehderi

Er. Marcus, 1965

Gymnodoris sp. 6

Cyerce katiae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Costasiella sp. 4

Placida barackobamai

McCarthy, Krug & Á. Valdés, 2017

Reticulidia fungia

Brunckhorst & Gosliner, 1993

Niparaya regiscorona

(Bertsch, 1972)

Odontoglaja guamensis

Rudman, 1978

Odontoglaja sp. 1

Philinopsis speciosa

Pease, 1860

Gastropteron minutum

Ong & Gosliner, 2017

Sagaminopteron sp. 1

Ilbia mariana

Hoff & Carlson, 1990

Elysia sp. 5

Elysia sp.12

Kaloplocamus acutus

Baba, 1949

Aegires malinus

Fahey & Gosliner, 2004

Aegires villosus

Farran, 1905

Miamira sinuata

(van Hasselt, 1824)

Chromodoris magnifica

(Quoy & Gaimard, 1832)

Glossodoris rufomarginata

(Bergh, 1890)

Hypselodoris infucata

(Rüppell & Leuckart, 1830)

Thorunna australis

(Risbec, 1928)

Phyllidia elegans

Bergh, 1869

Dermatobranchus rodmani

Gosliner & Fahey, 2011

Janolus mirabilis

Baba & Abe, 1970

Coryphellina delicata

(Gosliner & Willan, 1991)

Coryphellina sp. 2

Favorinus japonicus

Baba, 1949

Phestilla minor

Rudman, 1981

Nakamigawaia nakanoae

Hellem & Malaquias, 2022

Goniobranchus sp. 5

Tenellia sp.25

Sagaminopteron multimaculatum

Ong & Gosliner, 2017

Mariaglaja tsurugensis

(Baba & Abe, 1959)

Halgerda carlsoni

Rudman, 1978

Colpodaspis thompsoni

G. H. Brown, 1979

Diniatys dentifer

(A. Adams, 1850)

Sagaminopteron nigropunctatum

Carlson & Hoff, 1973

Siphopteron nigromarginatum

Gosliner, 1989

Plakobranchus ocellatus

van Hasselt, 1824

Thuridilla carlsoni

Gosliner, 1995

Thuridilla kathae

Gosliner, 1995

Thuridilla livida

(Baba, 1955)

Costasiella kuroshimae

M. Ichikawa, 1993

Stiliger sp. 1

Pleurobranchus forskalii

Rüppell & Leuckart, 1828

Hexabranchus lacer

(Cuvier, 1804)

Polycera sp. 1

Kaloplocamus dokte

Vallès & Gosliner, 2006

Nembrotha livingstonei

J. K. Allan, 1933

Ceratodoris nakamotoensis

(Hamatani, 2001)

Gymnodoris citrina

(Bergh, 1877)

Gymnodoris okinawae

Baba, 1936

Gymnodoris sp. 1

Gymnodoris suggens

(Risbec, 1928)

Gymnodoris sp. 7

Gymnodoris tuberculosa

Knutson & Gosliner, 2014

Miamira miamirana

(Bergh, 1875)

Chromodoris annae

Bergh, 1877

Chromodoris strigata

Rudman, 1982

Hypselodoris bullockii

(Collingwood, 1881)

Diversidoris crocea

(Rudman, 1986)

Verconia varians

(Pease, 1871)

Thorunna furtiva

Bergh, 1878

Phyllidiella annulata

(J. E. Gray, 1853)

Phyllidiella granulata

Brunckhorst, 1993

Costasiella sp. 5

Goniobranchus sp. 3

Phyllodesmium sp. 1

Gastropteron sp. 4

Doridoidea spp.

Facelinidae sp. 2

Limapontiidae spp.

Vellicolla amphorella

(A. Adams, 1862)

Micromelo guamensis

(Quoy & Gaimard, 1825)

Nembrothinae spp.

Hantazuia imagawai

Korshunova, Fletcher & Martynov, 2025

Hantazuia sp. 1

Thuridilla albopustulosa

Gosliner, 1995

Phanerophthalmus luteus

(Quoy & Gaimard, 1833)

Chelidonura hirundinina

(Quoy & Gaimard, 1833)

Chelidonura pallida

Risbec, 1951

Notarchus indicus

Schweigger, 1820

Lobiger viridis

Pease, 1863

Elysia thompsoni

K. R. Jensen, 1993

Cyerce kikutarobabai

Hamatani, 1976

Cyerce blackburnae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Costasiella sp. 2

Plocamopherus pecoso

Valles & Gosliner, 2006

Gymnodoris sp. 3

Gymnodoris sp. 5

Ardeadoris egretta

Rudman, 1984

Goniobranchus reticulatus

(Quoy & Gaimard, 1832)

Goniobranchus verrieri

(Crosse, 1875)

Mexichromis pusilla

(Bergh, 1874)

Glossodoris hikuerensis

(Pruvot-Fol, 1954)

Glossodoris buko

S. B. Matsuda & Gosliner, 2018

Hypselodoris whitei

(A. Adams & Reeve, 1850)

Ardeadoris angustolutea

(Rudman, 1990)

Diversidoris flava

(Eliot, 1904)

Thorunna halourga

R. F. Johnson & Gosliner, 2001

Dermatobranchus albus

(Eliot, 1904)

Coryphellina exoptata

(Gosliner & Willan, 1991)

Favorinus mirabilis

Baba, 1955

Favorinus tsuruganus

Baba & Abe, 1964

Phyllodesmium macphersonae

(Burn, 1962)

Limenandra fusiformis

(Baba, 1949)

Baeolidia japonica

Baba, 1933

Phestilla lugubris

(Bergh, 1870)

Doriprismatica sp. 1

Micromelo barbarae

Feliciano, Malaquias, Stout, Brenzinger, Gosliner & Á. Valdés, 2021

Cerberilla sp. 6

Niparaya sp. 2

Phyllidiidae spp.

Tenellia sp.45

Aeolidina sp. 8

Pruvotfolia sp. 4

Siphopteron sp. 3

Tenellia sp. 7

Elysia sp.29

Godiva sp. 4

Vellicolla muscaria

(A. A. Gould, 1859)

Halgerda sp. 3

Tenellia sp.76

Niparaya sp. 8

Abronica payaso

A. Y. Kim & Gosliner, 2024

Phanerophthalmus anettae

Austin, Gosliner & Malaquias, 2018

Sakuraeolis marhe

Fernández-Simón & Moles, 2023

Dermatobranchus sp. 5

Chromodoris dianae

Gosliner & Behrens, 1998

Aplysia sp. 1

Thuridilla bayeri

(Er. Marcus, 1965)

Gymnodoris nagoensis

De Souza-Canal, R. Nakano & Á. Valdés, 2025

Facelina sp. 8

Crimora sp. 1

Jorunna rubescens

(Bergh, 1876)

Tritoniopsis elegans

(Audouin, 1826)

Tubulophilinopsis pilsbryi

(Eliot, 1900)

Tubulophilinopsis gardineri

(Eliot, 1903)

Atys semistriatus

Pease, 1860

Spinoaglaja orientalis

(Baba, 1949)

Elysia marginata

(Pease, 1871)

Elysia pusilla

(Bergh, 1872)

Elysia rufescens

(Pease, 1871)

Elysia sp. 1

Elysia sp. 7

Elysia tomentosa

K. R. Jensen, 1997

Thuridilla gracilis

(Risbec, 1928)

Thuridilla undula

Gosliner, 1995

Cyerce goodheartae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Berthellina delicata

(Pease, 1861)

Pleurobranchus albiguttatus

(Bergh, 1905)

Plocamopherus ceylonicus

(Kelaart, 1858)

Plocamopherus imperialis

Angas, 1864

Plocamopherus maculapodium

Valles & Gosliner, 2006

Nembrotha chamberlaini

Gosliner & Behrens, 1997

Nembrotha lineolata

Bergh, 1905

Nembrotha milleri

Gosliner & Behrens, 1997

Nembrotha sp. 1

Martadoris sp. 1

Bermudella sp. 1

Trapania euryeia

Gosliner & Fahey, 2008

Trapania naeva

Gosliner & Fahey, 2008

Aegires citrinus

Pruvot-Fol, 1930

Gymnodoris feyae

De Souza-Canal & Valdés, 2025

Doriopsis pecten

(Collingwood, 1881)

Carminodoris estrelyado

(Gosliner & Behrens, 1998)

Halgerda willeyi

Eliot, 1904

Ceratosoma sp. 1

Chromodoris aspersa

(A. A. Gould, 1852)

Chromodoris alcalai

Gosliner, 2021

Goniobranchus hintuanensis

(Gosliner & Behrens, 1998)

Goniobranchus rubrocornutus

(Rudman, 1985)

Mexichromis lemniscata

(Quoy & Gaimard, 1832)

Hypselodoris krakatoa

Gosliner & R. F. Johnson, 1999

Hypselodoris maculosa

(Pease, 1871)

Verconia alboannulata

(Rudman, 1986)

Mexichromis trilineata

(A. Adams & Reeve, 1850)

Dendrodoris tuberculosa

(Quoy & Gaimard, 1832)

Ceratophyllidia sp. 1

Phyllidia varicosa

Lamarck, 1801

Dermatobranchus fasciatus

Gosliner & Fahey, 2011

Longibranchus putnami

(Fernández-Simón & Moles, 2023)

Cratena sp. 2

Facelina sp. 1

Bulbaeolidia alba

(Risbec, 1928)

Abronica purpureoanulata

(Baba, 1961)

Hermaea sp. 1

Nembrotha sp. 3

Nembrotha sp. 4

Siphopteron sp. 5

Tenellia nakapila

A. Y. Kim & Gosliner, 2024

Hypselodoris variobranchia

Gosliner & R. F. Johnson, 2018

Favorinus sp. 1

Facelina sp. 2

Tenellia sp.22

Aeolidina spp.

Jorunna parva

(Baba, 1938)

Phyllodesmium sp. 2

Martadoris sp. 2

Hermaea sp. 3

Chromodorididae sp. 3

Gymnodoris sp.15

Gymnodoris sp.16

Tritoniidae sp. 1

Tenellia sp.37

Gymnodoris sp.19

Plocamopherus sp. 2

Favorinus sp. 5

Ercolania endophytophaga

K. R. Jensen, 1999

Phidiana sp. 2

Goniobranchus sp. 9

Stiliger sp.10

Polybranchia spp.

Noumeaella sp. 4

Thecacera sp.12

Polycera sp.12

Gymnodoris sp.27

Niparaya sp. 6

Niparaya sp. 9

Doriopsilla sp. 3

Gymnodoris sp.31

Bursatella ocelligera

(Bergh, 1902)

Thuridilla sp. 8

Trapania sp.18

Gymnodoris sp.34

Hypselodoris sp.19

Unknown Species

Syphonota geographica

(A. Adams & Reeve, 1850)

Tylodina sp. 1

Tomoberthella martensi

(Pilsbry, 1896)

Goniobranchus coi

(Risbec, 1956)

Jorunna sp. 1

Chelidonura varians

Eliot, 1903

Phyllodesmium briareum

(Bergh, 1896)

Bornella stellifera

(A. Adams & Reeve, 1848)

Colpodaspis sp. 1

Colpodaspis sp. 2

Haminoeidae sp. 2

Philine elegans

Bergh, 1905

Hermania orca

(Gosliner, 1988)

Mariaglaja inornata

(Baba, 1949)

Chelidonura sp. 1

Noalda sp. 1

Metaruncina setoensis

(Baba, 1954)

Aplysia oculifera

A. Adams & Reeve, 1850

Dolabrifera dolabrifera

(Rang, 1828)

Petalifera punctulata

(Tapparone Canefri, 1874)

Petalifera ramosa

Baba, 1959

Stylocheilus striatus

(Quoy & Gaimard, 1832)

Elysia amakusana

Baba, 1955

Elysia lobata

A. A. Gould, 1852

Thuridilla hoffae

Gosliner, 1995

Mourgona sp. 1

Sohgenia palauensis

Hamatani, 1991

Ercolania subviridis

(Baba, 1959)

Sacoproteus smaragdinus

(Baba, 1949)

Pleurobranchus grandis

Pease, 1868

Polycera sp. 7

Polycera risbeci

Odhner, 1941

Polycera sp. 2

Thecacera picta

Baba, 1972

Plocamopherus maculatus

(Pease, 1860)

Tambja sagamiana

(Baba, 1955)

Pelagella felis

(Baba, 1949)

Ceratodoris lambat

(Gosliner, 2004)

Trapania aurata

Rudman, 1987

Trapania darvelli

Rudman, 1987

Aegires exeches

Fahey & Gosliner, 2004

Gymnodoris ceylonica

(Kelaart, 1858)

Gymnodoris inornata

(Bergh, 1880)

Gymnodoris sp.33

Gymnodoris subornata

Baba, 1960

Aldisa zavorensis

Tibiriçá, Pola & Cervera, 2017

Doriopsis granulosa

Pease, 1860

Doris viridis

(Pease, 1861)

Carminodoris grandiflora

(Pease, 1860)

Discodoris boholiensis

Bergh, 1877

Halgerda elegans

Bergh, 1905

Halgerda onna

Fahey & Gosliner, 2001

Jorunna sp. 3

Sebadoris fragilis

(Alder & Hancock, 1864)

Sebadoris nubilosa

(Pease, 1871)

Hallaxa hileenae

Gosliner & S. Johnson, 1994

Hallaxa iju

Gosliner & S. Johnson, 1994

Hallaxa indecora

(Bergh, 1905)

Hallaxa paulinae

Gosliner & S. Johnson, 1994

Ceratosoma palliolatum

Rudman, 1988

Chromodoris sp.14

Goniobranchus decorus

(Pease, 1860)

Chromodoris lineolata

(van Hasselt, 1824)

Goniobranchus sinensis

(Rudman, 1985)

Hypselodoris apolegma

(Yonow, 2001)

Mexichromis multituberculata

(Baba, 1953)

Verconia sp. 2

Mexichromis aurora

(R. F. Johnson & Gosliner, 1998)

Thorunna purpuropedis

Rudman & S. Johnson, 1985

Dendrodoris elongata

Baba, 1936

Dendrodoris fumata

(Rüppell & Leuckart, 1830)

Dendrodoris sp. 1

Phyllidia babai

Brunckhorst, 1993

Phyllidia carlsonhoffi

Brunckhorst, 1993

Phyllidia exquisita

Brunckhorst, 1993

Phyllidia picta

Pruvot-Fol, 1957

Phyllidia scottjohnsoni

Brunckhorst, 1993

Phyllidiidae sp. 1

Phyllidiella nigra

(van Hasselt, 1824)

Phyllidiopsis fissurata

Brunckhorst, 1993

Phyllidiopsis krempfi

Pruvot-Fol, 1957

Phyllidiopsis sphingis

Brunckhorst, 1993

Janolus sp. 2

Lomanotus vermiformis

Eliot, 1908

Melibe engeli

Risbec, 1937

Melibe japonica

Eliot, 1913

Marianina rosea

(Pruvot-Fol, 1930)

Samla takashigei

Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017

Annulorhina mandapamensis

K. P. Rao, 1968

Cratena simba

Edmunds, 1970

Phidiana salaamica

Rudman, 1980

Phyllodesmium longicirrum

(Bergh, 1905)

Phyllodesmium magnum

Rudman, 1991

Phyllodesmium poindimiei

(Risbec, 1928)

Pruvotfolia sp. 3

Limenandra rosanae

Carmona, Pola, Gosliner & Cervera, 2014

Baeolidia moebii

Bergh, 1888

Cerberilla affinis

Bergh, 1888

Cerberilla annulata

(Quoy & Gaimard, 1832)

Pleurolidia juliae

Burn, 1966

Tenellia diversicolor

(Baba, 1975)

Tenellia ornata

(Baba, 1937)

Eubranchus sp.11

Phestilla melanobrachia

Bergh, 1874

Gymnodoris mariefranceae

De Souza-Canal & Valdés, 2025

Kaloplocamus sp. 1

Kaloplocamus sp. 2

Pelagella longicornis

Paz-Sedano, Smirnoff, Gosliner & Pola, 2023

Paradoris sp. 2

Pelagella sp. 4

Tenellia sp.27

Polycera sp. 5

Dermatobranchus spp.

Hypselodoris lacuna

Gosliner & R. F. Johnson, 2018

Thorunna sp. 2

Murphydoris adusta

Paz-Sedano, Smirnoff, Candás, Gosliner & Pola, 2022

Murphydoris maracabranchia

Paz-Sedano, Smirnoff, Candás, Gosliner & Pola, 2022

Tenellia sp.10

Hallaxa sp. 2

Gastropteridae sp. 3

Phyllodesmium acanthorhinum

E. Moore & Gosliner, 2014

Paradoris sp. 5

Cratena sp. 3

Trapania sp. 9

Bulbaeolidia japonica

(Eliot, 1913)

Ercolania sp. 2

Niparaya sp. 3

Pruvotfolia sp. 2

Abronica sp. 1

Tritoniopsis cf. frydis

Spinoaglaja sp. 1

Philinopsis sp. 2

Gymnodoris sp.14

Marionia spp.

Janolus sp.11

Baeolidia variabilis

Carmona, Pola, Gosliner & Cervera, 2014

Tenellia sp.77

Gymnodoris sp.17

Chromodoris sp. 4

Phanerophthalmus albotriangulatus

Austin, Gosliner & Malaquias, 2018

Gymnodoris sp.20

Favorinus sp. 8

Tenellia sp.55

Mourgona sp. 3

Phestilla sp. 3

Niparaya sp. 4

Armina occulta

Mehrotra, Caballer & Chavanich, 2017

Armina scotti

Mehrotra, Caballer & Chavanich, 2017

Ercolania sp. 4

Gymnodoris sp.22

Eubranchus sp.21

Tenellia sp.72

Abronica sp. 5

Unidentia sandramillenae

Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017

Plocamopherus sp. 4

Tenellia sp.23

Samla sp. 2

Pelagella sp. 8

Hermaea wrangeliae

(M. Ichikawa, 1993)

Facelinidae sp.11

Eubranchus sp.27

Eubranchus sp.28

Tritonia sp. 7

Discodorididae spp.

Dendrodoris sp. 4

Niparaya sp. 7

Elysia minima

M. Ichikawa, 1993

Polycera sp.15

Gymnodoris plebeia

(Bergh, 1877)

Unidentia spp.

Aglajidae sp. 5

Janolus sp.18

Gastropteridae spp.

Niparaya sp.10

Phyllidia sp. 3

Gymnodoris bicolor

(Alder & Hancock, 1864)

Niparaya sp.11

Ercolania sp. 8

Thuridilla sp. 6

Coryphellina spp.

Atys naucum

(Linnaeus, 1758)

Trapania tamaraw

Smirnoff, Donohoo & Gosliner, 2022

Jorunna spp.

Aglajidae sp. 8

Goniobranchus cf. variatus

(Risbec, 1928)

Elysia sp.36

Mourgona sp. 4

Favorinus sp.12

Gymnodoris wanaporum

De Souza-Canal & Valdés, 2025

Biuve sp. 1

Elysia sp.38

Chromodorididae sp. 9

Rostanga sp.13

Martadoris sp. 3

Catriona lucerna

Korshunova, Martynov, Lundin & Malmberg, 2022

Phyllodesmium sp. 8