Sea Slugs in Okinawa Island (Onna and Yomitan area) 513

Chromodoris willani

Rudman, 1982

Tomoberthella martensi

(Pilsbry, 1896)

Chromodoris colemani

Rudman, 1982

Reticulidia fungia

Brunckhorst & Gosliner, 1993

Ardeadoris egretta

Rudman, 1984

Goniobranchus kuniei

(Pruvot-Fol, 1930)

Chromodoris sp. 3

Phyllidiella pustulosa

(Cuvier, 1804)

Halgerda carlsoni

Rudman, 1978

Sagaminopteron psychedelicum

Carlson & Hoff, 1974

Aldisa albatrossae

Elwood, Á. Valdés & Gosliner, 2000

Halgerda diaphana

Fahey & Gosliner, 1999

Halgerda elegans

Bergh, 1905

Hypselodoris apolegma

(Yonow, 2001)

Phyllidia coelestis

Bergh, 1905

Notodoris citrina

Bergh, 1875

Phyllidia elegans

Bergh, 1869

Phyllidiopsis xishaensis

(G.-Y. Lin, 1983)

Samla bicolor

(Kelaart, 1858)

Ardeadoris cruenta

(Rudman, 1986)

Phyllidia willani

Brunckhorst, 1993

Cadlinella ornatissima

(Risbec, 1928)

Goniobranchus sp. 1

Hypselodoris bullockii

(Collingwood, 1881)

Hypselodoris maculosa

(Pease, 1871)

Phyllidiidae sp. 1

Goniobranchus sp. 3

Halgerda tessellata

(Bergh, 1880)

Halgerda brunneomaculata

Carlson & Hoff, 1993

Phyllidia picta

Pruvot-Fol, 1957

Phyllidiopsis sphingis

Brunckhorst, 1993

Lamprohaminoea cymbalum

(Quoy & Gaimard, 1833)

Bornella anguilla

S. Johnson, 1984

Chromodoris aspersa

(A. A. Gould, 1852)

Janolus sp. 2

Lamprohaminoea ovalis

(Pease, 1868)

Siphopteron tigrinum

Gosliner, 1989

Elysia mercieri

(Pruvot-Fol, 1930)

Pleurobranchus forskalii

Rüppell & Leuckart, 1828

Aegires lemoncello

Fahey & Gosliner, 2004

Ceratosoma sp. 1

Goniobranchus geometricus

(Risbec, 1928)

Goniobranchus reticulatus

(Quoy & Gaimard, 1832)

Hypselodoris sp.18

Verconia norba

(Er. Marcus & Ev. Marcus, 1970)

Verconia romeri

(Risbec, 1928)

Verconia simplex

(Pease, 1871)

Mariaglaja inornata

(Baba, 1949)

Sagaminopteron ornatum

Tokioka & Baba, 1964

Aplysia nigrocincta

E. von Martens, 1880

Pleurobranchus peronii

Cuvier, 1804

Paradoris sp. 1

Ardeadoris angustolutea

(Rudman, 1990)

Diversidoris crocea

(Rudman, 1986)

Thorunna furtiva

Bergh, 1878

Thorunna halourga

R. F. Johnson & Gosliner, 2001

Phyllidia babai

Brunckhorst, 1993

Phyllidia scottjohnsoni

Brunckhorst, 1993

Pleurehdera sp. 1

Nembrotha chamberlaini

Gosliner & Behrens, 1997

Jorunna funebris

(Kelaart, 1858)

Hallaxa indecora

(Bergh, 1905)

Chromodoris strigata

Rudman, 1982

Ardeadoris averni

(Rudman, 1985)

Glossodoris cf. cincta

(Bergh, 1888)

Glossodoris hikuerensis

(Pruvot-Fol, 1954)

Phyllidiopsis pipeki

Brunckhorst, 1993

Lamprohaminoea evelinae

Oskars & Malaquias, 2020

Thecacera pacifica

(Bergh, 1884)

Tritoniopsis elegans

(Audouin, 1826)

Hypselodoris purpureomaculosa

Hamatani, 1995

Siphopteron nigromarginatum

Gosliner, 1989

Elysia rufescens

(Pease, 1871)

Cyerce kikutarobabai

Hamatani, 1976

Costasiella paweli

M. Ichikawa, 1993

Costasiella sp. 2

Kaloplocamus dokte

Vallès & Gosliner, 2006

Aegires flores

Fahey & Gosliner, 2004

Halgerda onna

Fahey & Gosliner, 2001

Chromodoris sp.14

Goniobranchus setoensis

(Baba, 1938)

Glossodoris rufomarginata

(Bergh, 1890)

Ardeadoris symmetrica

(Rudman, 1990)

Diversidoris flava

(Eliot, 1904)

Thorunna purpuropedis

Rudman & S. Johnson, 1985

Hancockia sp. 2

Goniobranchus sp. 4

Phyllidiidae spp.

Phyllidiidae sp. 2

Gastropteron sp. 3

Elysia obtusa

Baba, 1938

Elysia asbecki

Wägele, Stemmer, Burghardt & Händeler, 2010

Costasiella usagi

M. Ichikawa, 1993

Gymnodoris okinawae

Baba, 1936

Halgerda albocristata

Gosliner & Fahey, 1998

Rostanga sp. 2

Miamira sinuata

(van Hasselt, 1824)

Goniobranchus decorus

(Pease, 1860)

Mexichromis pusilla

(Bergh, 1874)

Glossodoris sp. 1

Verconia sp. 4

Thorunna australis

(Risbec, 1928)

Phyllidia exquisita

Brunckhorst, 1993

Phyllidiopsis cardinalis

Bergh, 1876

Dermatobranchus albus

(Eliot, 1904)

Cratena sp. 1

Facelina bourailli

(Risbec, 1928)

Tenellia ornata

(Baba, 1937)

Volvatella viridis

Hamatani, 1976

Tenellia sp.97

Goniobranchus coi

(Risbec, 1956)

Siphopteron brunneomarginatum

(Carlson & Hoff, 1974)

Elysia lobata

A. A. Gould, 1852

Plakobranchus ocellatus

van Hasselt, 1824

Thuridilla livida

(Baba, 1955)

Thuridilla splendens

(Baba, 1949)

Roboastra gracilis

(Bergh, 1877)

Trapania naeva

Gosliner & Fahey, 2008

Gymnodoris feyae

De Souza-Canal & Valdés, 2025

Hallaxa hileenae

Gosliner & S. Johnson, 1994

Goniobranchus albopunctatus

Garrett, 1879

Chromodoris alcalai

Gosliner, 2021

Goniobranchus rufomaculatus

(Pease, 1871)

Glossodoris buko

S. B. Matsuda & Gosliner, 2018

Ardeadoris tomsmithi

(Bertsch & Gosliner, 1989)

Thorunna daniellae

(Kay & D. K. Young, 1969)

Phyllidiella annulata

(J. E. Gray, 1853)

Dermatobranchus rodmani

Gosliner & Fahey, 2011

Moridilla brockii

Bergh, 1888

Tenellia sp.11

Diaphorodoris sp. 2

Tenellia sp.10

Phyllodesmium acanthorhinum

E. Moore & Gosliner, 2014

Aeolidina spp.

Halgerda sp. 3

Tyrannodoris nikolasi

(Pola, Padula, Gosliner & Cervera, 2014)

Anteaeolidiella sp. 3

Hypselodoris sp.24

Thuridilla vataae

(Risbec, 1928)

Odontoglaja guamensis

Rudman, 1978

Dolabrifera dolabrifera

(Rang, 1828)

Stylocheilus striatus

(Quoy & Gaimard, 1832)

Elysia sp. 7

Kaloplocamus acutus

Baba, 1949

Bermudella sp. 1

Aegires villosus

Farran, 1905

Gymnodoris sp. 3

Aldisa zavorensis

Tibiriçá, Pola & Cervera, 2017

Platydoris inornata

Dorgan, Á. Valdés & Gosliner, 2002

Hallaxa paulinae

Gosliner & S. Johnson, 1994

Goniobranchus fidelis

(Kelaart, 1858)

Hypselodoris bollandi

Gosliner & R. F. Johnson, 1999

Hypselodoris rositoi

Gosliner & R. F. Johnson, 2018

Verconia varians

(Pease, 1871)

Thorunna florens

(Baba, 1949)

Phyllidia larryi

(Brunckhorst, 1993)

Phyllidiopsis krempfi

Pruvot-Fol, 1957

Dermatobranchus fortunatus

(Bergh, 1888)

Dermatobranchus semilunus

Gosliner & Fahey, 2011

Dermatobranchus fasciatus

Gosliner & Fahey, 2011

Coryphellina delicata

(Gosliner & Willan, 1991)

Phyllodesmium magnum

Rudman, 1991

Pruvotfolia sp. 3

Phestilla melanobrachia

Bergh, 1874

Paradoris sp. 4

Micromelo barbarae

Feliciano, Malaquias, Stout, Brenzinger, Gosliner & Á. Valdés, 2021

Janolus sp.13

Vellicolla muscaria

(A. A. Gould, 1859)

Glossodoris sp. 2

Facelinidae sp.11

Doriopsilla sp. 2

Gymnodoris sp.31

Tritonia bollandi

V. G. Smith & Gosliner, 2003

Jorunna rubescens

(Bergh, 1876)

Chelidonura amoena

Bergh, 1905

Odontoglaja sp. 1

Julia zebra

Kawaguchi, 1981

Bosellia sp. 1

Elysia sp. 8

Thuridilla flavomaculata

Gosliner, 1995

Thuridilla hoffae

Gosliner, 1995

Cyerce bourbonica

Yonow, 2012

Costasiella fridae

Fernández-Simón & Moles, 2023

Berthellina delicata

(Pease, 1861)

Polycera sp. 1

Aegires exeches

Fahey & Gosliner, 2004

Aldisa pikokai

Bertsch & S. Johnson, 1982

Platydoris cruenta

(Quoy & Gaimard, 1832)

Chromodoris annae

Bergh, 1877

Diversidoris aurantionodulosa

Rudman, 1987

Hypselodoris krakatoa

Gosliner & R. F. Johnson, 1999

Mexichromis mariei

(Crosse, 1872)

Verconia sp. 2

Mexichromis aurora

(R. F. Johnson & Gosliner, 1998)

Dendrodoris sp. 1

Ceratophyllidia sp. 1

Phyllidia ocellata

Cuvier, 1804

Phyllidiella granulata

Brunckhorst, 1993

Phyllidiopsis fissurata

Brunckhorst, 1993

Doto cervicenigra

Ortea & Bouchet, 1989

Cratena simba

Edmunds, 1970

Favorinus mirabilis

Baba, 1955

Godiva sp. 1

Bulbaeolidia alba

(Risbec, 1928)

Pleurolidia juliae

Burn, 1966

Tenellia diversicolor

(Baba, 1975)

Abronica purpureoanulata

(Baba, 1961)

Phestilla minor

Rudman, 1981

Verconia sp. 1

Thorunna sp. 4

Pelagella longicornis

Paz-Sedano, Smirnoff, Gosliner & Pola, 2023

Tenellia sp.51

Polycera sp. 5

Pteraeolidia semperi

(Bergh, 1870)

Facelina sp. 2

Phyllodesmium sp. 1

Hallaxa sp. 1

Dermatobranchus diagonalis

Gosliner & Fahey, 2011

Pruvotfolia sp. 1

Tritoniopsis cf. frydis

Doridoidea spp.

Baeolidia variabilis

Carmona, Pola, Gosliner & Cervera, 2014

Chromodorididae spp.

Tenellia sp.73

Chromodoris sp. 4

Halgerda labyrinthus

Donohoo & Gosliner, 2023

Murphydoris puncticulata

Paz-Sedano, Smirnoff, Candás, Gosliner & Pola, 2022

Costasiella sp. 4

Tenellia sp. 7

Phyllidia fontjei

Wägele, Raubold, Papu, Undap & Yonow, 2025

Hypselodoris sp.23

Aliculastrum cylindricum

(Helbling, 1779)

Spinoaglaja orientalis

(Baba, 1949)

Chelidonura hirundinina

(Quoy & Gaimard, 1833)

Metaruncina setoensis

(Baba, 1954)

Oxynoe viridis

(Pease, 1861)

Elysia pusilla

(Bergh, 1872)

Elysia sp. 1

Elysia flavipunctata

M. Ichikawa, 1993

Elysia sp. 5

Elysia thompsoni

K. R. Jensen, 1993

Thuridilla carlsoni

Gosliner, 1995

Thuridilla gracilis

(Risbec, 1928)

Thuridilla kathae

Gosliner, 1995

Mourgona sp. 1

Cyerce liliuokalaniae

K. Moreno, Gosliner, N. G. Wilson, Krug & Á. Valdés, 2025

Costasiella formicaria

(Baba, 1959)

Costasiella kuroshimae

M. Ichikawa, 1993

Sacoproteus smaragdinus

(Baba, 1949)

Hexabranchus lacer

(Cuvier, 1804)

Polycera japonica

Baba, 1949

Plocamopherus imperialis

Angas, 1864

Plocamopherus maculatus

(Pease, 1860)

Tambja morosa

(Bergh, 1877)

Martadoris sp. 1

Naisdoris liklik

(Gosliner, 2004)

Aegires incusus

Fahey & Gosliner, 2004

Aegires malinus

Fahey & Gosliner, 2004

Aegires petalis

Fahey & Gosliner, 2004

Doriopsis pecten

(Collingwood, 1881)

Carminodoris estrelyado

(Gosliner & Behrens, 1998)

Diaulula sp.

Tayuva lilacina

(A. A. Gould, 1852)

Platydoris sanguinea

Bergh, 1905

Rostanga sp. 1

Ceratosoma palliolatum

Rudman, 1988

Ceratosoma tenue

Abraham, 1876

Goniobranchus collingwoodi

(Rudman, 1987)

Goniobranchus hintuanensis

(Gosliner & Behrens, 1998)

Chromodoris lochi

Rudman, 1982

Chromodoris magnifica

(Quoy & Gaimard, 1832)

Chromodoris michaeli

Gosliner & Behrens, 1998

Goniobranchus roboi

(Gosliner & Behrens, 1998)

Goniobranchus sinensis

(Rudman, 1985)

Goniobranchus verrieri

(Crosse, 1875)

Mexichromis similaris

(Rudman, 1986)

Hypselodoris infucata

(Rüppell & Leuckart, 1830)

Hypselodoris whitei

(A. Adams & Reeve, 1850)

Hypselodoris zephyra

Gosliner & R. F. Johnson, 1999

Mexichromis multituberculata

(Baba, 1953)

Dendrodoris carbunculosa

(Kelaart, 1858)

Dendrodoris coronata

Kay & D. K. Young, 1969

Dendrodoris elongata

Baba, 1936

Phyllidia polkadotsa

Brunckhorst, 1993

Phyllidia varicosa

Lamarck, 1801

Phyllidiopsis annae

Brunckhorst, 1993

Phyllidiopsis shireenae

Brunckhorst, 1990

Madrella ferruginosa

Alder & Hancock, 1864

Melibe engeli

Risbec, 1937

Tritonia sp. 1

Samla riwo

(Gosliner & Willan, 1991)

Caloria indica

(Bergh, 1896)

Cratena sp.11

Cratena affinis

(Baba, 1949)

Favorinus japonicus

Baba, 1949

Phanerophthalmus lentigines

Austin, Gosliner & Malaquias, 2018

Cadlinella subornatissima

Baba, 1996

Nakamigawaia nakanoae

Hellem & Malaquias, 2022

Pelagella albopunctata

Paz-Sedano, Smirnoff, Gosliner & Pola, 2023

Tergiposacca longicerata

Cella, Carmona, Ekimova, Chichvarkhin, Schepetov & Gosliner, 2016

Pelagella sp. 3

Murphydoris sp. 4

Samla sp. 1

Hypselodoris variobranchia

Gosliner & R. F. Johnson, 2018

Haminoeidae sp. 3

Trapania sp. 4

Gymnodoris spp.

Hypselodoris lacuna

Gosliner & R. F. Johnson, 2018

Cratena sp. 3

Phyllodesmium undulatum

E. Moore & Gosliner, 2014

Niparaya sp. 2

Anteaeolidiella takanosimensis

(Baba, 1930)

Tritoniidae spp.

Tenellia sp.77

Facelinidae sp. 2

Tenellia sp. 4

Tenellia sp.37

Elysia sp.18

Facelina sp. 5

Polycera sp. 9

Ercolania endophytophaga

K. R. Jensen, 1999

Elysia sp.24

Berthella caledonica

(Risbec, 1928)

Phestilla viei

Mehrotra, Caballer & Chavanich, 2020

Baeolidia sp. 4

Eubranchus sp.36

Avaldesia spp.

Eubranchus sp.37

Chromodoris sp.13

Julia mishimaensis

Kawaguti & Yamasu, 1982

Edenttellina pseudochloris

(Kay, 1964)

Kaloplocamus sp. 6

Chromodorididae sp.10

Syphonota geographica

(A. Adams & Reeve, 1850)

Thuridilla albopustulosa

Gosliner, 1995

Diaphorodoris sp. 1

Tubulophilinopsis gardineri

(Eliot, 1903)

Phyllodesmium briareum

(Bergh, 1896)

Colpodaspis sp. 1

Diniatys dentifer

(A. Adams, 1850)

Haminoeidae sp. 2

Hermania orca

(Gosliner, 1988)

Chelidonura pallida

Risbec, 1951

Spinophallus falciphallus

(Gosliner, 2011)

Philinopsis speciosa

Pease, 1860

Siphopteron ladrones

(Carlson & Hoff, 1974)

Ilbia mariana

Hoff & Carlson, 1990

Aplysia argus

Rüppell & Leuckart, 1830

Aplysia oculifera

A. Adams & Reeve, 1850

Petalifera punctulata

(Tapparone Canefri, 1874)

Volvatella angeliniana

M. Ichikawa, 1993

Lobiger viridis

Pease, 1863

Elysia nealae

Ostergaard, 1955

Elysia sp. 2

Elysia tomentosa

K. R. Jensen, 1997

Elysia yaeyamana

Baba, 1936

Sohgenia palauensis

Hamatani, 1991

Costasiella sp. 1

Stiliger aureomarginatus

K. R. Jensen, 1993

Hermaea sp. 2

Kaloplocamus peludo

Vallès & Gosliner, 2006

Nembrotha livingstonei

J. K. Allan, 1933

Nembrotha sp. 1

Naisdoris sp. 5

Pelagella joubini

(Risbec, 1928)

Naisdoris sp. 1

Notodoris gardineri

Eliot, 1906

Notodoris serenae

Gosliner & Behrens, 1997

Gymnodoris citrina

(Bergh, 1877)

Gymnodoris sp. 1

Gymnodoris sp. 5

Gymnodoris sp.33

Asteronotus cespitosus

(van Hasselt, 1824)

Halgerda dalanghita

Fahey & Gosliner, 1999

Halgerda sp. 2

Platydoris scabra

(Cuvier, 1804)

Jorunna hervei

Innabi, Stout & Á. Valdés, 2023

Thordisa sp. 2

Discodorididae sp. 2

Hallaxa fuscescens

(Pease, 1871)

Miamira miamirana

(Bergh, 1875)

Miamira moloch

(Rudman, 1988)

Goniobranchus leopardus

(Rudman, 1987)

Doriprismatica atromarginata

(Cuvier, 1804)

Mexichromis lemniscata

(Quoy & Gaimard, 1832)

Ardeadoris electra

(Rudman, 1990)

Hypselodoris babai

Gosliner & Behrens, 2000

Hypselodoris emma

Rudman, 1977

Verconia alboannulata

(Rudman, 1986)

Mexichromis trilineata

(A. Adams & Reeve, 1850)

Hypselodoris tryoni

(Garrett, 1873)

Dendrodoris nigra

(W. Stimpson, 1855)

Dendrodoris tuberculosa

(Quoy & Gaimard, 1832)

Ceratophyllidia sp. 2

Dermatobranchus gonatophorus

van Hasselt, 1824

Dermatobranchus oculus

Gosliner & Fahey, 2011

Dermatobranchus striatus

van Hasselt, 1824

Janolus tricellariodes

Pola & Gosliner, 2019

Notobryon bijecurum

Baba, 1937

Notobryon wardi

Odhner, 1936

Marionia sp. 1

Cephalopyge trematoides

(Chun, 1889)

Coryphellina exoptata

(Gosliner & Willan, 1991)

Launsina tanyae

Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026

Coryphellina sp. 2

Herviella affinis

Baba, 1960

Herviella albida

Baba, 1966

Noumeaella isa

Ev. Marcus & Er. Marcus, 1970

Noumeaella rehderi

Er. Marcus, 1965

Phyllodesmium kabiranum

Baba, 1991

Phyllodesmium longicirrum

(Bergh, 1905)

Sakuraeolis kirembosa

Rudman, 1980

Aeolidiopsis ransoni

Pruvot-Fol, 1956

Tenellia beta

(Baba & Abe, 1964)

Tenellia sp. 2

Eubranchus sp.11

Siphopteron sp. 1

Costasiella sp. 5

Polycera sp. 3

Bornella sp. 1

Lomanotus sp. 1

Trapania tigger

Smirnoff, Donohoo & Gosliner, 2022

Doriprismatica sp. 1

Unidentia sp. 1

Paradoris sp. 2

Pelagella sp. 4

Tenellia nakapila

A. Y. Kim & Gosliner, 2024

Hydatina physis

(Linnaeus, 1758)

Dendrodoris sp. 2

Elysia spp.

Aegires sp. 1

Thorunna sp. 2

Thorunna sp. 3

Murphydoris sp. 3

Gastropteron sp. 4

Marionia sp. 2

Hallaxa sp. 2

Tenellia sp.22

Murphydoris cobbi

Paz-Sedano, Smirnoff, Candás, Gosliner & Pola, 2022

Ercolania sp. 2

Avaldesia albomacula

(J. M. Chan & Gosliner, 2007)

Eubranchus sp. 9

Elysia sp.10

Pelagella sp. 6

Coryphellina sp. 1

Janolus sp. 7

Atagema sp. 2

Cavolinia uncinata

(A. d'Orbigny, 1835)

Peltodoris murrea

(Abraham, 1877)

Janolus sp. 9

Hermaea sp. 3

Janolus sp.12

Favorinus sp. 2

Chromodorididae sp. 3

Phyllidia sp. 1

Elysia nigropunctata

(Pease, 1871)

Gymnodoris sp.21

Favorinus sp. 8

Mourgona sp. 3

Limapontiidae spp.

Anteaeolidiella sp. 1

Discodorididae sp. 8

Tenellia sp.43

Ceratosoma bicolor

Baba, 1949

Tenellia sp.72

Mariaglaja tsurugensis

(Baba & Abe, 1959)

Noumeaella sp. 4

Godiva sp. 4

Marionia sp. 8

Glossodoris sp. 3

Samla sp. 2

Pelagella sp. 8

Rostanga sp. 9

Costasiella sp. 6

Tenellia sp.78

Verconia laboutei

(Rudman, 1986)

Elysia sp.33

Abronica payaso

A. Y. Kim & Gosliner, 2024

Phanerophthalmus anettae

Austin, Gosliner & Malaquias, 2018

Tenellia sp.40

Doridoidea sp. 6

Baeolidia rieae

Carmona, Pola, Gosliner & Cervera, 2014

Aplysia sp. 1

Verconia sp. 6

Tenellia sp.24

Gymnodoris bicolor

(Alder & Hancock, 1864)

Bursatella ocelligera

(Bergh, 1902)

Elysia sp.35

Thuridilla bayeri

(Er. Marcus, 1965)

Atys naucum

(Linnaeus, 1758)

Discodorididae sp.15

Facelinidae sp.14

Abronica turon

A. Y. Kim & Gosliner, 2024

Hexabranchus sanguineus

(Rüppell & Leuckart, 1830)

Goniodoridella sp.16

Janolus sp.20

Goniobranchus cf. variatus

(Risbec, 1928)

Hantazuia imagawai

Korshunova, Fletcher & Martynov, 2025

Coryphellina pseudolotos

Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022

Gymnodoris bricei

De Souza-Canal & Valdés, 2025

Discodorididae sp.25

Discodorididae sp.29

Haminoeidae sp.11

Gymnodoris sp.43

Elysia sp.40

Facelina sp. 8

Catriona lucerna

Korshunova, Martynov, Lundin & Malmberg, 2022

Phyllodesmium sp. 8

Crimora sp. 1

Dolabrifera sp. 1

Facelinidae sp.16

Trapania sp.23

Hypselodoris sp.22