Sea Slugs in Amakusa 470

Liloa porcellana

(A. A. Gould, 1859)

Phyllodesmium crypticum

Rudman, 1981

Armina japonica

(Eliot, 1913)

Hypselodoris festiva

(A. Adams, 1861)

Tritoniopsis elegans

(Audouin, 1826)

Marionia sp. 8

Atys semistriatus

Pease, 1860

Stiliger sp. 3

Goniodoridella borealis

Martynov, Sanamyan & Korshunova, 2015

Gymnodoris spp.

Gymnodoris sp.23

Lamprohaminoea sp. 4

Philine orientalis

A. Adams, 1855

Elysia nigrocapitata

Baba, 1957

Thecacera picta

Baba, 1972

Nembrotha lorosae

Pola, Paz-Sedano, Martín, L. Warren, Noble & Martín-Hervás, 2026

Bermudella japonica

(Baba, 1949)

Jorunna parva

(Baba, 1938)

Doto purpurea

Baba, 1949

Elysia cf. japonica

Eliot, 1913

Pruvotfolia sp. 4

Acteocina decorata

(Pilsbry, 1904)

Acteocina sp. 2

Goniodoridella sp.17

Elysia marginata

(Pease, 1871)

Calliopaea pusilla

(Baba, 1968)

Diaphorodoris mitsuii

(Baba, 1938)

Chromodoris orientalis

Rudman, 1983

Goniobranchus sp.16

Gymnodoris amakusana

(Baba, 1996)

Phanerophthalmus sp. 3

Haloa japonica

(Pilsbry, 1895)

Philinopsis gigliolii

(Tapparone Canefri, 1874)

Aplysia argus

Rüppell & Leuckart, 1830

Aplysia nigrocincta

E. von Martens, 1880

Petalifera punctulata

(Tapparone Canefri, 1874)

Placida babai

Ev. Marcus, 1982

Kaloplocamus albopunctatus

J.C. Wei & L.F. Kong, 2023

Tyrannodoris sp. 1

Aldisa albatrossae

Elwood, Á. Valdés & Gosliner, 2000

Goniobranchus sinensis

(Rudman, 1985)

Dermatobranchus primus

Baba, 1976

Phyllodesmium macphersonae

(Burn, 1962)

Stiliger sp. 5

Melanochlamys fukudai

S. Cooke, Hanson, Y. Hirano, Ornelas-Gatdula, Gosliner, A. V. Chernyshev & Á. Valdés, 2014

Haminoeidae sp. 7

Japonactaeon sp. 1

Acteocina exilis

(Dunker, 1860)

Retusa concentrica

(A. Adams, 1850)

Philinopsis speciosa

Pease, 1860

Oxynoe viridis

(Pease, 1861)

Stiliger berghi

Baba, 1937

Stiliger sp. 1

Hexabranchus lacer

(Cuvier, 1804)

Vayssierea felis

(Collingwood, 1881)

Pelagella castanea

(Alder & Hancock, 1845)

Ceratodoris hiroi

(Baba, 1938)

Gymnodoris inornata

(Bergh, 1880)

Aldisa zavorensis

Tibiriçá, Pola & Cervera, 2017

Hypselodoris sagamiensis

(Baba, 1949)

Eubranchus inabai

Baba, 1964

Acteocina sp.

Aeolidina spp.

Tenellia sp. 4

Discodorididae sp. 1

Trinchesiidae sp. 2

Phyllidiella pustulosa

(Cuvier, 1804)

Haminoeidae sp. 2

Hermania orca

(Gosliner, 1988)

Gastropteron bicornutum

Baba & Tokioka, 1965

Elysia sp.16

Thuridilla splendens

(Baba, 1949)

Stiliger sp. 2

Aplysiopsis minor

(Baba, 1959)

Hermaea noto

(Baba, 1959)

Tambja amakusana

Baba, 1987

Tambja morosa

(Bergh, 1877)

Tambja pulcherrima

Willan & Y.-W. Chang, 2017

Discodorididae sp. 3

Goniobranchus fidelis

(Kelaart, 1858)

Hypselodoris bullockii

(Collingwood, 1881)

Mexichromis mariei

(Crosse, 1872)

Verconia norba

(Er. Marcus & Ev. Marcus, 1970)

Verconia purpurea

(Baba, 1949)

Thorunna florens

(Baba, 1949)

Dendrodoris arborescens

(Collingwood, 1881)

Dendrodoris krusensternii

(J. E. Gray, 1850)

Doto pita

Er. Marcus, 1955

Sakuraeolis enosimensis

(Baba, 1930)

Baeolidia japonica

Baba, 1933

Baeolidia salaamica

(Rudman, 1982)

Tenellia ornata

(Baba, 1937)

Goniobranchus aureopurpureus

(Collingwood, 1881)

Miamira flavicostata

Baba, 1940

Facelina sp. 3

Tenellia sp.69

Pupa strigosa

(A. A. Gould, 1859)

Gymnodoris inariensis

Hamatani & Osumi, 2004

Tenellia sp.21

Favorinus sp. 9

Ceratosoma bicolor

Baba, 1949

Vellicolla muscaria

(A. A. Gould, 1859)

Leostyletus spp.

Aeolidina sp. 5

Plakobranchidae sp. 2

Gymnodoris boiteuxi

De Souza-Canal & Valdés, 2025

Thecacera pikachu

Pola, Paz-Sedano, Martín, L. Warren, Noble & Martín-Hervás, 2026

Runcinida elioti

(Baba, 1937)

Aplysia juliana

Quoy & Gaimard, 1832

Bursatella leachii

Blainville, 1817

Stylocheilus striatus

(Quoy & Gaimard, 1832)

Lobiger viridis

Pease, 1863

Elysia atroviridis

Baba, 1955

Elysia obtusa

Baba, 1938

Elysia tomentosa

K. R. Jensen, 1997

Stiliger aureomarginatus

K. R. Jensen, 1993

Pleurobranchaea japonica

Thiele, 1925

Pleurobranchus weberi

(Bergh, 1905)

Polycera hedgpethi

Er. Marcus, 1964

Tyrannodoris luteolineata

(Baba, 1936)

Ancula kariyana

Baba, 1990

Pelagella felis

(Baba, 1949)

Pelagella joubini

(Risbec, 1928)

Okenia echinata

Baba, 1949

Gymnodoris okinawae

Baba, 1936

Gymnodoris subornata

Baba, 1960

Doriopsis granulosa

Pease, 1860

Geitodoris lutea

Baba, 1937

Rostanga orientalis

Rudman & Avern, 1989

Jorunna sp. 3

Jorunna hervei

Innabi, Stout & Á. Valdés, 2023

Sclerodoris rubicunda

(Baba, 1949)

Goniobranchus leopardus

(Rudman, 1987)

Goniobranchus setoensis

(Baba, 1938)

Doriprismatica atromarginata

(Cuvier, 1804)

Ardeadoris averni

(Rudman, 1985)

Glossodoris buko

S. B. Matsuda & Gosliner, 2018

Glossodoris rufomarginata

(Bergh, 1890)

Hypselodoris maculosa

(Pease, 1871)

Hypselodoris maritima

(Baba, 1949)

Hypselodoris placida

(Baba, 1949)

Mexichromis multituberculata

(Baba, 1953)

Dermatobranchus ornatus

(Bergh, 1874)

Dermatobranchus semistriatus

Baba, 1949

Madrella ferruginosa

Alder & Hancock, 1864

Melibe papillosa

(de Filippi, 1867)

Doto bella

Baba, 1938

Kabeiro sp. 2

Samla takashigei

Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017

Leostyletus misakiensis

(Baba, 1960)

Eubranchus sp. 5

Favorinus pacificus

Baba, 1937

Favorinus tsuruganus

Baba & Abe, 1964

Phyllodesmium kabiranum

Baba, 1991

Phyllodesmium magnum

Rudman, 1991

Sakuraeolis sakuracea

Y. Hirano, 1999

Setoeolis inconspicua

(Baba, 1938)

Bulbaeolidia alba

(Risbec, 1928)

Limenandra fusiformis

(Baba, 1949)

Baeolidia moebii

Bergh, 1888

Cerberilla ambonensis

Bergh, 1905

Cerberilla sp. 1

Cerberilla sp. 2

Tenellia sibogae

(Bergh, 1905)

Phestilla melanobrachia

Bergh, 1874

Murphydoris sp. 4

Trapania sp. 4

Polycera abei

(Baba, 1960)

Atagema sp. 1

Tenellia sp. 8

Elysia sp.10

Tritoniidae spp.

Tritonia sp. 5

Favorinus sp. 2

Baeolidia variabilis

Carmona, Pola, Gosliner & Cervera, 2014

Gymnodoris sp.18

Cylichnatys angusta

(A. A. Gould, 1859)

Geitodoris sp. 1

Limapontiidae spp.

Tenellia futairo

(Baba, 1963)

Tenellia sp.23

Eubranchus sp.25

Hermaea sp. 4

Lomanotus sp. 8

Cavoliniidae spp.

Ringicula kurodai

Takeyama, 1935

Baeolidia sp. 4

Runcinida sp. 3

Acteocina gordonis

(Yokoyama, 1927)

Coryphellina spp.

Mnestia girardi

(Audouin, 1826)

Decorifer insignis

(Pilsbry, 1904)

Gymnodoris sp.36

Japonactaeon nipponensis

(Yamakawa, 1911)

Vellicolla ooformis

(Habe, 1952)

Elysia sp.37

Tambja gabrielae

Pola, Cervera & Gosliner, 2005

Thuridilla albopustulosa

Gosliner, 1995

Thuridilla vataae

(Risbec, 1928)

Siphopteron flavum

(Tokioka & Baba, 1964)

Bornella stellifera

(A. Adams & Reeve, 1848)

Retusa succincta

(A. Adams, 1862)

Niparaya regiscorona

(Bertsch, 1972)

Chelidonura amoena

Bergh, 1905

Biuve fulvipunctata

(Baba, 1938)

Chelidonura hirundinina

(Quoy & Gaimard, 1833)

Tubulophilinopsis lineolata

(H. Adams & A. Adams, 1854)

Sagaminopteron ornatum

Tokioka & Baba, 1964

Sagaminopteron pohnpei

(Hoff & Carlson, 1983)

Aplysia kurodai

(Baba, 1937)

Aplysia oculifera

A. Adams & Reeve, 1850

Dolabella auricularia

([Lightfoot], 1786)

Dolabrifera dolabrifera

(Rang, 1828)

Elysia amakusana

Baba, 1955

Elysia trisinuata

Baba, 1949

Costasiella formicaria

(Baba, 1959)

Ercolania boodleae

(Baba, 1938)

Pleurehdera sp. 1

Palio amakusana

Baba, 1960

Polycera sp. 1

Polycera sp. 2

Thecacera pennigera

(Montagu, 1813)

Kalinga ornata

Alder & Hancock, 1864

Naisdoris sp. 5

Ceratodoris barnardi

(Baba, 1937)

Trapania miltabrancha

Gosliner & Fahey, 2008

Idaliadoris sp. 1

Aegires citrinus

Pruvot-Fol, 1930

Gymnodoris alba

(Bergh, 1877)

Gymnodoris citrina

(Bergh, 1877)

Gymnodoris impudica

(Rüppell & Leuckart, 1830)

Atagema intecta

(Kelaart, 1858)

Carminodoris armata

Baba, 1993

Homoiodoris japonica

Bergh, 1882

Jorunna funebris

(Kelaart, 1858)

Jorunna cf. pantherina

(Angas, 1864)

Sebadoris fragilis

(Alder & Hancock, 1864)

Thordisa luteola

J. M. Chan & Gosliner, 2007

Hallaxa cryptica

Gosliner & S. Johnson, 1994

Cadlinella ornatissima

(Risbec, 1928)

Ceratosoma trilobatum

(J. E. Gray, 1850)

Goniobranchus albonares

(Rudman, 1990)

Goniobranchus albopunctatus

Garrett, 1879

Goniobranchus collingwoodi

(Rudman, 1987)

Goniobranchus roboi

(Gosliner & Behrens, 1998)

Hypselodoris apolegma

(Yonow, 2001)

Hypselodoris decorata

(Risbec, 1928)

Hypselodoris whitei

(A. Adams & Reeve, 1850)

Verconia nivalis

(Baba, 1937)

Verconia subnivalis

(Baba, 1987)

Hypselodoris tryoni

(Garrett, 1873)

Dendrodoris fumata

(Rüppell & Leuckart, 1830)

Dendrodoris guttata

(Odhner, 1917)

Dendrodoris tuberculosa

(Quoy & Gaimard, 1832)

Phyllidiopsis krempfi

Pruvot-Fol, 1957

Armina babai

(S. Tchang, 1934)

Armina comta

(Bergh, 1880)

Armina semperi

(Bergh, 1866)

Dermatobranchus otome

Baba, 1992

Armina rubida

(A. A. Gould, 1852)

Dermatobranchus striatellus

Baba, 1949

Janolus indicus

(Eliot, 1909)

Melibe japonica

Eliot, 1913

Melibe viridis

(Kelaart, 1858)

Marionia sp. 1

Tritonia sp. 1

Doto cervicenigra

Ortea & Bouchet, 1989

Coryphellina lotos

Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017

Caloria indica

(Bergh, 1896)

Cratena affinis

(Baba, 1949)

Facelina bilineata

Y. Hirano & M. Ito, 1998

Facelina quadrilineata

(Baba, 1930)

Phyllodesmium serratum

(Baba, 1949)

Cerberilla asamusiensis

Baba, 1940

Spurilla braziliana

MacFarland, 1909

Tenellia diversicolor

(Baba, 1975)

Tenellia pupillae

(Baba, 1961)

Abronica purpureoanulata

(Baba, 1961)

Phestilla lugubris

(Bergh, 1870)

Phanerophthalmus lentigines

Austin, Gosliner & Malaquias, 2018

Siphopteron makisig

Ong & Gosliner, 2017

Samla sp. 1

Tenellia nakapila

A. Y. Kim & Gosliner, 2024

Hypselodoris variobranchia

Gosliner & R. F. Johnson, 2018

Tenellia bughaw

A. Y. Kim & Gosliner, 2024

Polycera sp. 4

Elysia spp.

Abronica sp. 2

Trapania sp. 9

Melanochlamys ezoensis

(Baba, 1957)

Baeolidia sp. 1

Aliculastrum debile

(Pease, 1860)

Janolus sp. 6

Unidentia sp. 2

Noumeaella sp. 3

Limenandra confusa

Carmona, Pola, Gosliner & Cervera, 2014

Peltodoris murrea

(Abraham, 1877)

Catriona osezakiensis

Martynov, Korshunova, Lundin, Malmberg, 2022

Thecacera sp. 7

Tenellia sp.67

Tenellia sp.26

Philine sp. 1

Aeolidina sp.15

Stiliger sp. 8

Tenellia sp.72

Gymnodoris sp.25

Phestilla goniophaga

Hu, Zhang, Yiu, Xie & Qiu, 2020

Coryphellina sp. 4

Tenellia sp.19

Lomanotus sp. 7

Favorinus sp.10

Janolus sp.18

Aplysia japonica

G. B. Sowerby II, 1869

Plakobranchidae sp. 1

Gymnodoris sp.31

Ringicula doliaris

A. A. Gould, 1860

Sakuraeolis sp. 4

Pyrunculus phialus

(A. Adams, 1862)

Punctacteon fabreanus

(Crosse, 1873)

Pneumoderma sp. 1

Anteaeolidiella decorus

S.-Q. Zhang & S.-P. Zhang, 2023

Chromodorididae sp. 9

Cylichna biplicata

(A. Adams, 1850)

Gymnodoris sp.41

Unknown Species

Thecacera pacifica

(Bergh, 1884)

Goniobranchus coi

(Risbec, 1956)

Pleurobranchaea brockii

Bergh, 1897

Colpodaspis thompsoni

G. H. Brown, 1979

Diniatys dentifer

(A. Adams, 1850)

Phanerophthalmus perpallidus

Risbec, 1928

Spinoaglaja orientalis

(Baba, 1949)

Philine elegans

Bergh, 1905

Mariaglaja inornata

(Baba, 1949)

Gastropteron minutum

Ong & Gosliner, 2017

Siphopteron fuscum

(Baba & Tokioka, 1965)

Siphopteron nigromarginatum

Gosliner, 1989

Metaruncina setoensis

(Baba, 1954)

Aplysia gigantea

G. B. Sowerby II, 1869

Elysia asbecki

Wägele, Stemmer, Burghardt & Händeler, 2010

Elysia sp. 7

Cyerce kikutarobabai

Hamatani, 1976

Placida kevinleei

McCarthy, Krug & Á. Valdés, 2017

Berthellina delicata

(Pease, 1861)

Pleurobranchus forskalii

Rüppell & Leuckart, 1828

Pleurobranchus peronii

Cuvier, 1804

Polycera fujitai

Baba, 1937

Polycera japonica

Baba, 1949

Crimora lutea

Baba, 1949

Limacia ornata

(Baba, 1937)

Plocamopherus tilesii

Bergh, 1877

Nembrotha lineolata

Bergh, 1905

Roboastra gracilis

(Bergh, 1877)

Tambja sagamiana

(Baba, 1955)

Okenia babai

Hamatani, 1961

Ceratodoris pilosa

(Bouchet & Ortea, 1983)

Okenia rhinorma

Rudman, 2007

Trapania euryeia

Gosliner & Fahey, 2008

Aegires exeches

Fahey & Gosliner, 2004

Aegires villosus

Farran, 1905

Gymnodoris suggens

(Risbec, 1928)

Aldisa cooperi

Robilliard & Baba, 1972

Doris immonda

(Risbec, 1928)

Atagema spongiosa

(Kelaart, 1858)

Discodoris cebuensis

Bergh, 1877

Tayuva lilacina

(A. A. Gould, 1852)

Discodoris pardalis

(Alder & Hancock, 1864)

Paradoris tsurugensis

Baba, 1986

Rostanga risbeci

Baba, 1991

Rostanga sp. 2

Doriopsilla sp. 1

Thordisa sanguinea

Baba, 1955

Actinocyclus papillatus

(Bergh, 1878)

Hallaxa indecora

(Bergh, 1905)

Ceratosoma tenue

Abraham, 1876

Chromodoris aspersa

(A. A. Gould, 1852)

Goniobranchus geometricus

(Risbec, 1928)

Goniobranchus tumuliferus

(Collingwood, 1881)

Goniobranchus verrieri

(Crosse, 1875)

Hypselodoris emma

Rudman, 1977

Hypselodoris kaname

Baba, 1994

Hypselodoris zephyra

Gosliner & R. F. Johnson, 1999

Verconia varians

(Pease, 1871)

Dendrodoris nigra

(W. Stimpson, 1855)

Doriopsilla miniata

(Alder & Hancock, 1864)

Phyllidia ocellata

Cuvier, 1804

Phyllidia picta

Pruvot-Fol, 1957

Phyllidia varicosa

Lamarck, 1801

Phyllidiopsis annae

Brunckhorst, 1993

Armina magna

Baba, 1955

Dermatobranchus cymatilis

Gosliner & Fahey, 2011

Madrella gloriosa

Baba, 1949

Doto japonica

Odhner, 1936

Embletonia gracilis

Risbec, 1928

Samla bicolor

(Kelaart, 1858)

Coryphellina exoptata

(Gosliner & Willan, 1991)

Samla riwo

(Gosliner & Willan, 1991)

Eubranchopsis virginalis

Baba, 1949

Eubranchus mimeticus

Baba, 1975

Eubranchus sp. 1

Longibranchus putnami

(Fernández-Simón & Moles, 2023)

Myja hyotan

Martynov, Mehrotra, Chavanich, R. Nakano, Kashio, Lundin, Picton & Korshunova, 2019

Antonietta janthina

Baba & Hamatani, 1977

Cratena lineata

(Eliot, 1905)

Cratena simba

Edmunds, 1970

Favorinus japonicus

Baba, 1949

Phyllodesmium opalescens

Rudman, 1991

Cerberilla albopunctata

Baba, 1976

Pleurolidia juliae

Burn, 1966

Tenellia anulata

(Baba, 1949)

Tenellia sp. 5

Polycera sp. 3

Akera soluta

(Gmelin, 1791)

Cerberilla sp. 5

Nembrotha sp. 3

Atys sp. 1

Arminidae spp.

Cerberilla sp. 4

Gymnodoris sp.11

Micromelo barbarae

Feliciano, Malaquias, Stout, Brenzinger, Gosliner & Á. Valdés, 2021

Dermatobranchus spp.

Novaeolidiella janae

Carmona, Martín-Hervás, Pola, Gosliner & Cervera, 2026

Ceratodoris sp. 2

Bornella hermanni

Angas, 1864

Janolus sp. 5

Anteaeolidiella takanosimensis

(Baba, 1930)

Tenellia puti

A. Y. Kim & Gosliner, 2024

Marionia spp.

Bulla sp. 1

Doridoidea spp.

Ardeadoris sp. 1

Creseis acicula

(Rang, 1828)

Eubranchus sp.17

Acteon sieboldii

(Reeve, 1842)

Eubranchus cf. leopoldoi

Caballer, Ortea & Espinosa, 2001

Tenellia sp.33

Elysia sp.23

Murphydoris puncticulata

Paz-Sedano, Smirnoff, Candás, Gosliner & Pola, 2022

Eubranchus sp.18

Nihonbranchus horii

(Baba, 1960)

Placida daguilarensis

K. Jensen, 1990

Runcinida sp. 2

Eubranchus sp.24

Sakuraeolis sp. 2

Trinchesiidae sp. 1

Pseudobornella orientalis

Baba, 1932

Aeolidina sp. 3

Gymnodoris sp.27

Eubranchus sp.26

Tenellia sp.15

Cerberilla longibranchus

(Volodchenko, 1941)

Discodorididae spp.

Verconia laboutei

(Rudman, 1986)

Tenellia sp.40

Placida barackobamai

McCarthy, Krug & Á. Valdés, 2017

Janolus sp.17

Sakuraeolis marhe

Fernández-Simón & Moles, 2023

Haminoeidae sp. 8

Niparaya sp.10

Aplysia sp. 3

Tenellia sp.89

Eubranchus sp.33

Retusa spp.

Bursatella ocelligera

(Bergh, 1902)

Kaloplocamus japonicus

(Bergh, 1880)

Goniodoridella geminae

Paz-Sedano, Ekimova, Smirnoff, Gosliner & Pola, 2023

Gymnodoris fassioae

De Souza-Canal & Valdés, 2025

Tenellia sp.84

Pyrunculus tokyoensis

Habe, 1950

Punctacteon amakusaensis

T. Habe, 1961